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Acritarch

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Acritarchs (Template:Etymology)<ref>Template:Cite book</ref> are organic microfossils, known from the Precambrian to the present. The classification is a catch all term used to refer to any organic microfossils that cannot be assigned to other groups. Their diversity reflects major ecological events such as the appearance of predation and the Cambrian explosion. Many acritarchs likely represent resting cysts of single-celled marine phytoplanktonic algae, similar to those produced by living dinoflagellates.<ref>Template:Cite journal</ref>

DefinitionEdit

Acritarchs were originally defined as non-acid soluble (i.e. non-carbonate, non-siliceous) organic-walled microfossils consisting of a central cavity, and whose biological affinities cannot be determined with certainty.<ref>Template:Cite journal</ref><ref>Template:Cite journal</ref><ref>Template:Cite journal</ref> Most commonly they are composed of thermally altered acid insoluble carbon compounds (kerogen).

Acritarchs may include the remains of a wide range of quite different kinds of organisms—ranging from the egg cases of small metazoans to resting cysts of many kinds of chlorophyta (green algae). It is likely that most acritarch species from the Paleozoic represent various stages of the life cycle of algae that were ancestral to the dinoflagellates.<ref>Template:Cite journal</ref> The nature of the organisms associated with Precambrian acritarchs is generally not well understood, though many are probably related to unicellular marine algae. In theory, when the biological source (taxon) of an acritarch does become known, that particular microfossil is removed from the acritarchs and classified with its proper group.Template:Citation needed

While the classification of acritarchs into form genera is entirely artificial, it is not without merit, as the form taxa show traits similar to those of genuine taxa—for example an 'explosion' in the Cambrian and a mass extinction at the end of the Permian.Template:Citation needed

ClassificationEdit

The informal group, Acritarcha Evitt 1963, was originally divided into the Subgroups: Acanthomorphitae, Polygonomorphitae, Prismatomorphitae, Oömorphitae, Netromorphitae, Dinetromorphitae, Stephanomorphitae, Pteromorphitae, Herkomorphitae, Platymorphitae, Sphaeromorphitae, and Disphaeromorphitae.<ref>Template:Cite journal</ref><ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref><ref>Template:Cite journal</ref>

Acritarchs were most likely eukaryotes. While archaea and bacteria (prokaryotes) usually produce simple fossils of a very small size, eukaryotic unicellular fossils are usually larger and more complex, with external morphological projections and ornamentation such as spines and hairs that only eukaryotes can produce; as most acritarchs have external projections (e.g., hair, spines, thick cell membranes, etc.), they are predominantly eukaryotes, although simple eukaryote acritarchs also exist.<ref>Template:Cite journal</ref>

The recent application of atomic force microscopy, confocal microscopy, Raman spectroscopy, and other sophisticated analytic techniques to the study of the ultrastructure, life history, and systematic affinities of mineralized, but originally organic-walled microfossils,<ref>Template:Cite journal</ref><ref>Template:Cite journal</ref><ref>Template:Cite journal</ref><ref>Template:Cite journal</ref><ref>Template:Cite journal</ref> has shown that some acritarchs are actually fossilized microalgae. In the end, it may well be, as Moczydłowska et al. suggested in 2011, that many acritarchs will, in fact, turn out to be algae.<ref>Template:Cite journal</ref><ref name=Chamberlain2016>Template:Cite journal File:CC-BY icon.svg Material was copied from this source, which is available under a Creative Commons Attribution 4.0 International License.</ref>

OccurrenceEdit

Acritarchs are found in sedimentary rocks from the present back into the Archean.<ref>Template:Cite journal</ref> They are typically isolated from siliciclastic sedimentary rocks using hydrofluoric acid but are occasionally extracted from carbonate-rich rocks. They are excellent candidates for index fossils used for dating rock formations in the Paleozoic Era and when other fossils are not available. Because most acritarchs are thought to be marine (pre-Triassic), they are also useful for palaeoenvironmental interpretation.

The Archean and earliest Proterozoic microfossils termed "acritarchs" may actually be prokaryotes. The earliest eukaryotic acritarchs known (Template:As of) are from between 1950 and 2150 million years ago.<ref>Template:Cite journal</ref>

DiversityEdit

At about 1 billion years ago the organisms responsible for acritarchs started to increase in abundance, diversity, size, complexity of shape, and especially, the size and number of spines. Their populations crashed during periods of extensive worldwide glaciations that covered the majority of the planet, but they proliferated in the Cambrian explosion and reached their highest diversity in the Paleozoic. The increased spininess 1000 million years ago possibly resulted from the need for defense against predators, especially predators large enough to swallow them or tear them apart. Other groups of small organisms from the Neoproterozoic era also show signs of anti-predator defenses.<ref name="Bengtson2002OriginsOfPredation">Template:Cite encyclopedia</ref>

Further evidence that acritarchs were subject to herbivory around this time comes from a consideration of taxon longevity. The abundance of planktonic organisms that evolved between 1,700 and 1,400 million years ago was limited by nutrient availability – a situation which limits the origination of new species because the existing organisms are so specialised to their niches, and no other niches are available for occupation. Approximately 1,000 million years ago, species longevity fell sharply, suggesting that predation pressure, probably by protist herbivores, became an important factor. Predation would have kept populations in check, meaning that some nutrients were left unused, and new niches were available for new species to occupy.<ref name=Stanley2008>Template:Cite journal</ref>

EtymologyEdit

Acritarch was coined in 1963 from the Greek ákritos meaning confused (a kritēs, without critic) and archē meaning origin (confer archaic).<ref>definition of acritarch at dictionary.com</ref>

Selected generaEdit

There are over 900 described genera; the following is a partial list, from Fossilid.info.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>

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See alsoEdit

ReferencesEdit

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External linksEdit

|CitationClass=web }} Commission Internationale de Microflore du Paléozoique (CIMP), international commission for Palaeozoic palynology.

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|CitationClass=web }} The American Association of Stratigraphic Palynologists (AASP)

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