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Butterflyfish

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Template:Short description Template:Hatnote group Template:Automatic taxobox

The butterflyfish are a group of conspicuous tropical marine fish of the family Chaetodontidae; the bannerfish and coralfish are also included in this group. The approximately 129 species in 12 genera<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> are found mostly on the reefs of the Atlantic, Indian, and Pacific Oceans. A number of species pairs occur in the Indian and Pacific Oceans, members of the huge genus Chaetodon.

Butterflyfish look like smaller versions of angelfish (Pomacanthidae), but unlike these, lack preopercle spines at the gill covers. Some members of the genus Heniochus resemble the Moorish idol (Zanclus cornutus) of the monotypic Zanclidae. Among the paraphyletic Perciformes, the former are probably not too distantly related to butterflyfish, whereas the Zanclidae seem far less close.

Description and ecologyEdit

File:Schooling bannerfish.JPG
A school of false Moorish idols, Heniochus diphreutes

Butterflyfish mostly range from Template:Convert in length. The largest species, the lined butterflyfish and the saddle butterflyfish, C. ephippium, grow to Template:Convert. The common name references the brightly coloured and strikingly patterned bodies of many species, bearing shades of black, white, blue, red, orange, and yellow. Other species are dull in colour. Butterflyfish are a boundless, different group of marine percoids with delegates on practically all coral reef frameworks and in every single tropical ocean. Their bright and color patterns have drawn in much consideration, creating an abundance of data about their conduct and environment.<ref>Smith, W. (2003). The evolution of the laterophysic connection with a revised phylogeny and taxonomy of butterflyfishes (Teleostei: Chaetodontidae). Cladistics the International Journal of the Willi Hennig Society., 19(4), 287–306. https://doi.org/10.1016/S0748-3007(03)00070-7</ref> Many have eyespots on their flanks and dark bands across their eyes, not unlike the patterns seen on butterfly wings.<ref name="EoF"> Template:Cite book</ref> Their deep, laterally narrow bodies are easily noticed through the profusion of reef life. The conspicuous coloration of butterflyfish may be intended for interspecies communication. Butterflyfish have uninterrupted dorsal fins with tail fins that may be rounded or truncated, but are never forked.

Generally diurnal and frequenting waters less than Template:Convert deep (though some species descend to Template:Convert, butterflyfish stick to particular home ranges. These corallivores are especially territorial, forming pairs and staking claim to a specific coral head. Contrastingly, the zooplankton feeders form large conspecific groups. By night, butterflyfish hide in reef crevices and exhibit markedly different coloration.

Their coloration also makes them popular aquarium fish. However, most species feed on coral polyps and sea anemones. Balancing the relative populations of prey and predator is complex, leading hobby aquarists to focus on the few generalists and specialist zooplankton feeders.

Butterflyfish are pelagic spawners; that is, they release many buoyant eggs into the water, which become part of the plankton, floating with the currents until hatching. The fry go through a tholichthys stage, wherein the body of the postlarval fish is covered in large, bony plates extending from the head. They lose their bony plates as they mature.<ref name="EoF" /> Only one other family of fish, the scats (Scatophagidae) express such an armored stage.

Taxonomy, systematics and evolutionEdit

The Chaetodontidae can be, but are not usually, divided into two lineages that arguably are subfamilies. The subfamily name Chaetodontinae is a little-used leftover from the period when the Pomacanthidae and Chaetodontidae were united under the latter name as a single family. Hence, Chaetodontinae is today considered a junior synonym of Chaetodontidae. In any case, one lineage of Chaetodontidae (in the modern sense) contains the "typical" butterflyfish around Chaetodon, while the other unites the bannerfish and coralfish genera. As the Perciformes are highly paraphyletic, the precise relationships of the Chaetodontidae as a whole are badly resolved.<ref name="fw2007">Fessler & Westneat (2007)</ref>

Chaetodontidae is classified within the suborder Percoidei by the 5th edition of Fishes of the World, but they are placed in an unnamed clade which sits outside the superfamily Percoidea. This clade contains 7 families which appear to have some relationship to Acanthuroidei, Monodactylidae, and Priacanthidae.<ref name = Nelson5>Template:Cite book</ref> Other authorities have paced the family in the order Chaetodontiformes alongside the family Leiognathidae.<ref name="Betancur-Rodriguez-4">Template:Cite journal</ref> Presently, based on the most recent phylogenetic analyses, Eschmeyer's Catalog of Fishes places it in the large, diverse order Acanthuriformes.<ref name=":1">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> Molecular analyses suggest that the Chaetodontidae shared a common ancestor with the Scatophagidae and Pomacanthidae that lived during the Early Eocene (54 Ma).<ref name=":0">Template:Cite journal</ref>

Before DNA sequencing, the taxonomy was confused about whether to treat these as species or subspecies. Also, numerous subgenera have been proposed for splitting out of Chaetodon, and it is becoming clear how to subdivide the genus if that is desired.<ref>Fessler & Westneat (2007), Hsu et al. (2007)</ref>

Fossil recordEdit

The fossil record of this group is disputed and marginal. Their restriction to coral reefs means their carcasses are liable to be dispersed by scavengers, overgrown by corals, and any that do fossilize will not long survive erosion. Many Eocene-aged fish such as Pygaeus have been variously referred to Chaetodontidae in the past; if used for calibration, it can be deduced that most living genera were probably distinct by the end of the Paleogene 23 Mya.<ref>Sepkoski (2002), Fessler & Westneat (2007)</ref> However, these Eocene genera cannot be reliably referred to Chaetodontidae.<ref name=":0" />

The earliest confirmed fossil specimens of Chaetodontidae are an indeterminate larval specimen from Frauenweiler, Germany, two tholichthys specimens from the Menilite Formation of Poland, as well as the extinct species Chaetodon (Blumchaetodon) wattsi from Italy; all date to the Early Oligocene. The occurrence of the first confirmed butterflyfish in the Early Oligocene is supported by molecular phylogenies indicating that the family may have arose during the Late Eocene.<ref name=":0" />

GeneraEdit

The bannerfish-coralfish lineage can be further divided in two groups; these might be considered tribes, but have not been formally named. Genera are listed in order of the presumed phylogeny, from the most ancient to the youngest:<ref name="fw2007" /><ref>{{#invoke:Cite taxon|main|fishbase|genus=|species=|subspecies=}}</ref>

Bannerfish/coralfish lineage 1:

Bannerfish/coralfish lineage 2:

The "typical" butterflyfishes may eventually come to contain more genera; see Chaetodon:

TimelineEdit

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 id:ANK 	 value:rgb(0.4,0.3,0.196)
 id:HER	  value:teal
 id:HAD	  value:green
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 id:white        value:white
 id:cenozoic     value:rgb(0.54,0.54,0.258)
 id:paleogene     value:rgb(0.99,0.6,0.32) 
 id:paleocene     value:rgb(0.99,0.65,0.37) 
 id:eocene     value:rgb(0.99,0.71,0.42) 
 id:oligocene     value:rgb(0.99,0.75,0.48) 
 id:neogene     value:rgb(0.999999,0.9,0.1) 
 id:miocene     value:rgb(0.999999,0.999999,0) 
 id:pliocene     value:rgb(0.97,0.98,0.68)  
 id:quaternary   value:rgb(0.98,0.98,0.5)
 id:pleistocene   value:rgb(0.999999,0.95,0.68)
 id:holocene   value:rgb(0.999,0.95,0.88)

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bar:NAM2

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align:center textcolor:black fontsize:M mark:(line,black) width:25 
shift:(7,-4)

bar:periodtop
from: -66   till:  -55.8    color:paleocene  text:Paleocene
from: -55.8   till:  -33.9    color:eocene  text:Eocene 
from: -33.9   till:  -23.03    color:oligocene  text:Oligocene            
from: -23.03    till: -5.332    color:miocene    text:Miocene
from: -5.332    till: -2.588    color:pliocene    text:Plio.
from: -2.588    till: -0.0117   color:pleistocene    text:Pleist.
from: -0.0117    till: 0    color:holocene    text:H.

bar:eratop
from: -66   till:  -23.03    color:paleogene  text:Paleogene         
from: -23.03    till: -2.588    color:neogene    text:Neogene
from: -2.588    till: 0   color:quaternary    text:Q.

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align:left fontsize:M mark:(line,white) width:5 anchor:till align:left

color:oligocene bar:NAM1  from:	-28.4	till:	0	text:	Chaetodon
color:miocene bar:NAM2  from:	-23.03	till:	0	text:	Chelmon

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align:center textcolor:black fontsize:M mark:(line,black) width:25

bar:period
from: -66   till:  -55.8    color:paleocene  text:Paleocene
from: -55.8   till:  -33.9    color:eocene  text:Eocene 
from: -33.9   till:  -23.03    color:oligocene  text:Oligocene            
from: -23.03    till: -5.332    color:miocene    text:Miocene
from: -5.332    till: -2.588    color:pliocene    text:Plio.
from: -2.588    till: -0.0117   color:pleistocene    text:Pleist.
from: -0.0117    till: 0    color:holocene    text:H.

bar:era
from: -66   till:  -23.03    color:paleogene  text:Paleogene         
from: -23.03    till: -2.588    color:neogene    text:Neogene
from: -2.588    till: 0   color:quaternary    text:Q.

</timeline>

GalleryEdit

Further readingEdit

  • Pratchett, Morgan S. & Berumen, Michael L. & Kapoor, B.G. [Editors] : Biology of Butterflyfishes. CRC Press, 2014. Template:ISBN

ReferencesEdit

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