Timeline of the evolutionary history of life

Template:Short description Template:Distinguish Template:For Template:Life timeline The timeline of the evolutionary history of life represents the current scientific theory outlining the major events during the development of life on planet Earth. Dates in this article are consensus estimates based on scientific evidence, mainly fossils.

In biology, evolution is any change across successive generations in the heritable characteristics of biological populations. Evolutionary processes give rise to diversity at every level of biological organization, from kingdoms to species, and individual organisms and molecules, such as DNA and proteins. The similarities between all present day organisms imply a common ancestor from which all known species, living and extinct, have diverged. More than 99 percent of all species that ever lived (over five billion)<ref name="Book-Biology">Template:Harvnb</ref> are estimated to be extinct.<ref name="StearnsStearns2000">Template:Cite book</ref><ref name="NYT-20141108-MJN">Template:Cite newsTemplate:Cbignore</ref> Estimates on the number of Earth's current species range from 10 million to 14 million,<ref name="MillerSpoolman2012">Template:Harvnb</ref> with about 1.2 million or 14% documented, the rest not yet described.<ref name="PLoS-20110823">Template:Cite journal</ref> However, a 2016 report estimates an additional 1 trillion microbial species, with only 0.001% described.<ref name="NSF-2016002">Template:Cite news</ref>

There has been controversy between more traditional views of steadily increasing biodiversity, and a newer view of cycles of annihilation and diversification, so that certain past times, such as the Cambrian explosion, experienced maximums of diversity followed by sharp winnowing.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref><ref>Template:Harvnb Four diagrams of evolutionary models</ref>

Extinction

{{#invoke:Labelled list hatnote|labelledList|Main article|Main articles|Main page|Main pages}}

File:Geological time spiral.png

Visual representation of the history of life on Earth as a spiral

Species go extinct constantly as environments change, as organisms compete for environmental niches, and as genetic mutation leads to the rise of new species from older ones. At long irregular intervals, Earth's biosphere suffers a catastrophic die-off, a mass extinction,<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> often comprising an accumulation of smaller extinction events over a relatively brief period.<ref name="bbc.co.uk">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>

The first known mass extinction was the Great Oxidation Event 2.4 billion years ago, which killed most of the planet's obligate anaerobes. Researchers have identified five other major extinction events in Earth's history, with estimated losses below:<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>

• End Ordovician: 440 million years ago, 86% of all species lost, including most<ref name= Mitchell>Template:Cite journal</ref> graptolites
• Late Devonian: 375 million years ago, 75% of species lost, including most trilobites
• End Permian, The Great Dying: 251 million years ago, 96% of species lost, including tabulate corals, and most trees and synapsids
• End Triassic: 200 million years ago, 80% of species lost, including all conodonts
• End Cretaceous: 66 million years ago, 76% of species lost, including all ammonites, mosasaurs, plesiosaurs, pterosaurs, and nonavian dinosaurs

Smaller extinction events have occurred in the periods between, with some dividing geologic time periods and epochs. The Holocene extinction event is currently under way.<ref name="pnas98_1"> Template:Cite journal </ref>

Factors in mass extinctions include continental drift, changes in atmospheric and marine chemistry, volcanism and other aspects of mountain formation, changes in glaciation, changes in sea level, and impact events.<ref name="bbc.co.uk"/>

Detailed timeline

In this timeline, Ma (for megaannum) means "million years ago," ka (for kiloannum) means "thousand years ago," and ya means "years ago."

Hadean Eon

File:FullMoon2010.jpg

Moon

{{#invoke:Labelled list hatnote|labelledList|Main article|Main articles|Main page|Main pages}} 4540 Ma – 4031 Ma

Date	Event
4540 Ma	Planet Earth forms from the accretion disc revolving around the young Sun, perhaps preceded by formation of organic compounds necessary for life in the surrounding protoplanetary disk of cosmic dust.<ref name="Space-20120329">Template:Cite news</ref><ref>Template:Cite journal</ref>
4510 Ma	According to the giant-impact hypothesis, the Moon originated when Earth and the hypothesized planet Theia collided, sending into orbit myriad moonlets which eventually coalesced into our single Moon.<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref><ref>Template:Cite journal</ref> The Moon's gravitational pull stabilised Earth's fluctuating axis of rotation, setting up regular climatic conditions favoring abiogenesis.<ref>Template:Cite journal</ref>
4404 Ma	Evidence of the first liquid water on Earth which were found in the oldest known zircon crystals.<ref>Template:Cite journal</ref>
4280–3770 Ma	Earliest possible appearance of life on Earth.<ref name="NAT-20170301">Template:Cite journal</ref><ref name="NYT-20170301">Template:Cite newsTemplate:Cbignore</ref><ref name="BBC-20170301">{{#invoke:citation/CS1|citation	CitationClass=web }} </ref><ref name="4.3b oldest"> Template:Cite news </ref>

Archean Eon

{{#invoke:Labelled list hatnote|labelledList|Main article|Main articles|Main page|Main pages}}

File:Acasta gneiss.jpg

Fragment of the Acasta Gneiss exhibited at the Museum of Natural History in Vienna

File:Cyanobacterial-algal mat.jpg

The cyanobacterial-algal mat, salty lake on the White Sea seaside

File:Halobacteria.jpg

Halobacterium sp. strain NRC-1

4031 Ma – 2500 Ma

Date	Event
4100 Ma	citation	CitationClass=web }}</ref><ref>Template:Cite journal</ref>
4100–3800 Ma	Late Heavy Bombardment (LHB): extended barrage by meteoroids impacting the inner planets. Thermal flux from widespread hydrothermal activity during the LHB may have aided abiogenesis and life's early diversification.<ref>Template:Cite journal</ref> Possible remains of biotic life were found in 4.1 billion-year-old rocks in Western Australia.<ref name="AP-20151019">Template:Cite news</ref><ref name="PNAS-20151014-pdf">Template:Cite journal</ref>
4000 Ma	Formation of a greenstone belt of the Acasta Gneiss of the Slave craton in northwest Canada - the oldest known rock belt.<ref name="Bjornerud">Template:Harvnb</ref>
3900–2500 Ma	Cells resembling prokaryotes appear.<ref>Template:Cite journal</ref> These first organisms are believed to have been chemoautotrophs, using carbon dioxide as a carbon source and oxidizing inorganic materials to extract energy.
3800 Ma	Formation of a greenstone belt of the Isua complex in western Greenland, whose isotope frequencies suggest the presence of life.<ref name="Bjornerud" /> The earliest evidence for life on Earth includes: 3.8 billion-year-old biogenic hematite in a banded iron formation of the Nuvvuagittuq Greenstone Belt in Canada;<ref>Template:Cite news</ref> graphite in 3.7 billion-year-old metasedimentary rocks in western Greenland;<ref name="NG-20131208">Template:Cite journal</ref> and microbial mat fossils in 3.48 billion-year-old sandstone in Western Australia.<ref name="AP-20131113">Template:Cite news</ref><ref name="AST-20131108">Template:Cite journal</ref>
3800–3500 Ma	Last universal common ancestor (LUCA):<ref>Template:Cite journal</ref><ref>Template:Cite journal</ref> split between bacteria and archaea.<ref>Template:Cite journal</ref> Bacteria develop primitive photosynthesis, which at first did not produce oxygen.<ref>Template:Cite journal</ref> These organisms exploit a proton gradient to generate adenosine triphosphate (ATP), a mechanism used by virtually all subsequent organisms.<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref><ref>Template:Cite journal</ref><ref>Template:Cite journal</ref>
3000 Ma	Photosynthesizing cyanobacteria using water as a reducing agent and producing oxygen as a waste product.<ref name="Buick, R. 2008">Template:Cite journal</ref> Free oxygen initially oxidizes dissolved iron in the oceans, creating iron ore. Oxygen concentration in the atmosphere slowly rises, poisoning many bacteria and eventually triggering the Great Oxygenation Event.
2800 Ma	Oldest evidence for microbial life on land in the form of organic matter-rich paleosols, ephemeral ponds and alluvial sequences, some bearing microfossils.<ref name="Beraldi-Campesi">Template:Cite journal</ref>

Proterozoic Eon

File:Endomembrane system diagram en (edit).svg

Detail of the eukaryote endomembrane system and its components

File:Mikrofoto.de-Blepharisma japonicum 15.jpg

Blepharisma japonicum, a free-living ciliated protozoan

File:DickinsoniaCostata.jpg

Dickinsonia costata, an iconic Ediacaran organism, displays the characteristic quilted appearance of Ediacaran enigmata.

{{#invoke:Labelled list hatnote|labelledList|Main article|Main articles|Main page|Main pages}} 2500 Ma – 539 Ma. Contains the Palaeoproterozoic, Mesoproterozoic and Neoproterozoic eras.

Date	Event
2500 Ma	Great Oxidation Event led by cyanobacteria's oxygenic photosynthesis.<ref name="Buick, R. 2008" /> Commencement of plate tectonics with old marine crust dense enough to subduct.<ref name="Bjornerud" />
2400 Ma	Possible land fungi evidence from molecules.
2023 Ma	Formation of the Vredefort impact structure, one of the largest and oldest verified impact structures on Earth. The crater is estimated to have been between Template:Convert across when it first formed.<ref>Template:Cite journal</ref>
By 1850 Ma	Eukaryotic cells, containing membrane-bound organelles with diverse functions, probably derived from prokaryotes engulfing each other via phagocytosis. (See Symbiogenesis and Endosymbiont). Bacterial viruses (bacteriophages) emerge before or soon after the divergence of the prokaryotic and eukaryotic lineages.<ref>Template:Cite journal</ref> Red beds show an oxidising atmosphere, favouring the spread of eukaryotic life.<ref>Template:Harvnb</ref><ref>Template:Cite journal</ref><ref>Template:Cite journal</ref>
1500 Ma	Volyn biota, a collection of exceptionally well-preserved microfossils with varying morphologies.<ref name=fra2022>Template:Cite journal</ref>
1300 Ma	Earliest land fungi.<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
By 1200 Ma	Meiosis and sexual reproduction in single-celled eukaryotes, possibly even in the common ancestor of all eukaryotes<ref>Template:Harvnb</ref> or in the RNA world.<ref>Template:Cite journal</ref> Sexual reproduction may have increased the rate of evolution.<ref name="dateref">Template:Cite journal</ref>
By 1000 Ma	First non-marine eukaryotes move onto land. They were photosynthetic and multicellular, indicating that plants evolved much earlier than originally thought.<ref>Template:Cite journal</ref>
750 Ma	Beginning of animal evolution.<ref name="NYT-20191127">Template:Cite newsTemplate:Cbignore</ref><ref name="BE-20161205">Template:Cite journal</ref>
720–630 Ma	Possible global glaciation<ref name="Hoffman1998">Template:Cite journal</ref><ref>Template:Harvnb</ref> which increased the atmospheric oxygen and decreased carbon dioxide, and was either caused by land plant evolution<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref> or resulted in it.<ref name="Zarsky-et-al-2022">Template:Cite journal</ref> Opinion is divided on whether it increased or decreased biodiversity or the rate of evolution.<ref>Template:Cite journal</ref><ref>Template:Cite journal</ref><ref name="Corsetti2006">Template:Cite journal</ref>
600 Ma	Accumulation of atmospheric oxygen allows the formation of an ozone layer.<ref name="Formation of the Ozone Layer">{{#invoke:citation/CS1|citation	CitationClass=web }}</ref> Previous land-based life would probably have required other chemicals to attenuate ultraviolet radiation.<ref name="Beraldi-Campesi" />
580–542 Ma	Ediacaran biota, the first large, complex aquatic multicellular organisms.<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
580–500 Ma	Cambrian explosion: most modern animal phyla appear.<ref name="BerkeleyCambrian">{{#invoke:citation/CS1|citation	CitationClass=web }}</ref><ref name="BristolUCEtiming">{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
550–540 Ma	Ctenophora (comb jellies),<ref>Template:Cite journal</ref> Porifera (sponges),<ref>Template:Cite journal</ref> Anthozoa (corals and sea anemones),<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref> Ikaria wariootia (an early Bilaterian).<ref>Template:Cite journal</ref>

Phanerozoic Eon

{{#invoke:Labelled list hatnote|labelledList|Main article|Main articles|Main page|Main pages}} 539 Ma – present

The Phanerozoic Eon (Greek: period of well-displayed life) marks the appearance in the fossil record of abundant, shell-forming and/or trace-making organisms. It is subdivided into three eras, the Paleozoic, Mesozoic and Cenozoic, with major mass extinctions at division points.

Palaeozoic Era

{{#invoke:Labelled list hatnote|labelledList|Main article|Main articles|Main page|Main pages}} Template:More citations needed 538.8 Ma – 251.9 Ma and contains the Cambrian, Ordovician, Silurian, Devonian, Carboniferous and Permian periods.

File:Nautilus belauensis profile.jpg

With only a handful of species surviving today, the Nautiloids flourished during the early Paleozoic era, from the Late Cambrian, where they constituted the main predatory animals.<ref>Template:Cite journal</ref>

File:Haikouichthys 3d.png

CitationClass=web }}</ref>

File:Sa-fern.jpg

CitationClass=web }}</ref>

File:Dimetrodon grandis 3D Model Reconstruction.png

Synapsids such as Dimetrodon were the largest terrestrial vertebrates in the Permian period, 299 to 251 million years ago.

Date	Event
535 Ma	Major diversification of living things in the oceans: arthropods (e.g. trilobites, crustaceans), chordates, echinoderms, molluscs, brachiopods, foraminifers and radiolarians, etc.
530 Ma	The first known footprints on land date to 530 Ma.<ref>Template:Cite journal</ref>
520 Ma	Earliest graptolites.<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
511 Ma	Earliest crustaceans.<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
505 Ma	Fossilization of the Burgess Shale
500 Ma	Jellyfish have existed since at least this time.
485 Ma	First vertebrates with true bones (jawless fishes).
450 Ma	First complete conodonts and echinoids appear.
440 Ma	First agnathan fishes: Heterostraci, Galeaspida, and Pituriaspida.
420 Ma	Earliest ray-finned fishes, trigonotarbid arachnids, and land scorpions.<ref name="Garwood">Template:Cite journal</ref>
410 Ma	First signs of teeth in fish. Earliest Nautilida, lycophytes, and trimerophytes.
488–400 Ma	First cephalopods (nautiloids)<ref>Template:Cite journal</ref> and chitons.<ref>Template:Cite journal</ref>
395 Ma	First lichens, stoneworts. Earliest harvestmen, mites, hexapods (springtails) and ammonoids. The earliest known tracks on land named the Zachelmie trackways which are possibly related to icthyostegalians.<ref>Template:Cite journal</ref>
375 Ma	Tiktaalik, a lobe-finned fish with some anatomical features similar to early tetrapods. It has been suggested to be a transitional species between fish and tetrapods.<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
365 Ma	Acanthostega is one of the earliest vertebrates capable of walking.<ref>Template:Cite journal</ref>
363 Ma	By the start of the Carboniferous Period, the Earth begins to resemble its present state. Insects roamed the land and would soon take to the skies; sharks swam the oceans as top predators,<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref> and vegetation covered the land, with seed-bearing plants and forests soon to flourish. Four-limbed tetrapods gradually gain adaptations which will help them occupy a terrestrial life-habit.
360 Ma	First crabs and ferns. Land flora dominated by seed ferns. The Xinhang forest grows around this time.<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
350 Ma	First large sharks, ratfishes, and hagfish; first crown tetrapods (with five digits and no fins and scales).
350 Ma	Diversification of amphibians.<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
325-335 Ma	First Reptiliomorpha.<ref>Template:Cite journal</ref>
330-320 Ma	First amniote vertebrates (Paleothyris).<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
320 Ma	Synapsids (precursors to mammals) separate from sauropsids (reptiles) in late Carboniferous.<ref name="Amniota">{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
305 Ma	The Carboniferous rainforest collapse occurs, causing a minor extinction event, as well as paving the way for amniotes to become dominant over amphibians and seed plants over ferns and lycophytes. First diapsid reptiles (e.g. Petrolacosaurus).
280 Ma	Earliest beetles, seed plants and conifers diversify while lepidodendrids and sphenopsids decrease. Terrestrial temnospondyl amphibians and pelycosaurs (e.g. Dimetrodon) diversify in species.
275 Ma	Therapsid synapsids separate from pelycosaur synapsids.
265 Ma	Gorgonopsians appear in the fossil record.<ref>Template:Cite journal</ref>
251.9–251.4 Ma	The Permian–Triassic extinction event eliminates over 90-95% of marine species. Terrestrial organisms were not as seriously affected as the marine biota. This "clearing of the slate" may have led to an ensuing diversification, but life on land took 30 million years to completely recover.<ref name="SahneyBenton2008RecoveryFromProfoundExtinction">Template:Cite journal</ref>

Mesozoic Era

{{#invoke:Labelled list hatnote|labelledList|Main article|Main articles|Main page|Main pages}} Template:More citations needed

File:Utatsusaurus BW.jpg

Utatsusaurus is the earliest-known ichthyopterygian.

File:Plateosaurus panorama.jpg

Plateosaurus engelhardti

File:Cycas circinalis.jpg

Cycas circinalis

File:Tyrannosaurus Rex Jane.jpg

For about 150 million years, dinosaurs were the dominant land animals on Earth.

From 251.9 Ma to 66 Ma and containing the Triassic, Jurassic and Cretaceous periods.

Date	Event
250 Ma	Mesozoic marine revolution begins: increasingly well adapted and diverse predators stress sessile marine groups; the "balance of power" in the oceans shifts dramatically as some groups of prey adapt more rapidly and effectively than others.
250 Ma	Triadobatrachus massinoti is the earliest known frog.
248 Ma	Sturgeon and paddlefish (Acipenseridae) first appear.
245 Ma	Earliest ichthyosaurs
240 Ma	Increase in diversity of cynodonts and rhynchosaurs
225 Ma	Earliest dinosaurs (prosauropods), first cardiid bivalves, diversity in cycads, bennettitaleans, and conifers. First teleost fishes. First mammals (Adelobasileus).
220 Ma	Seed-producing Gymnosperm forests dominate the land; herbivores grow to huge sizes to accommodate the large guts necessary to digest the nutrient-poor plants.Template:Citation needed First flies and turtles (Odontochelys). First coelophysoid dinosaurs. First mammals from small-sized cynodonts, which transitioned towards a nocturnal, insectivorous, and endothermic lifestyle.
205 Ma	Massive Triassic/Jurassic extinction. It wipes out all pseudosuchians except crocodylomorphs, who transitioned to an aquatic habitat, while dinosaurs took over the land and pterosaurs filled the air.
200 Ma	First accepted evidence for viruses infecting eukaryotic cells (the group Geminiviridae).<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref> However, viruses are still poorly understood and may have arisen before "life" itself, or may be a more recent phenomenon. Major extinctions in terrestrial vertebrates and large amphibians. Earliest examples of armoured dinosaurs.
195 Ma	First pterosaurs with specialized feeding (Dorygnathus). First sauropod dinosaurs. Diversification in small, ornithischian dinosaurs: heterodontosaurids, fabrosaurids, and scelidosaurids.
190 Ma	Pliosauroids appear in the fossil record. First lepidopteran insects (Archaeolepis), hermit crabs, modern starfish, irregular echinoids, corbulid bivalves, and tubulipore bryozoans. Extensive development of sponge reefs.
176 Ma	First Stegosaurian dinosaurs.
170 Ma	Earliest salamanders, newts, cryptoclidids, elasmosaurid plesiosaurs, and cladotherian mammals. Sauropod dinosaurs diversify.
168 Ma	First lizards.
165 Ma	First rays and glycymeridid bivalves. First vampire squids.<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
163 Ma	Pterodactyloid pterosaurs first appear.<ref>Template:Cite news</ref>
161 Ma	Ceratopsian dinosaurs appear in the fossil record (Yinlong) and the oldest known eutherian mammal: Juramaia.
160 Ma	Multituberculate mammals (genus Rugosodon) appear in eastern China.
155 Ma	First blood-sucking insects (ceratopogonids), rudist bivalves, and cheilostome bryozoans. Archaeopteryx, a possible ancestor to the birds, appears in the fossil record, along with triconodontid and symmetrodont mammals. Diversity in stegosaurian and theropod dinosaurs.
131 Ma	First pine trees.
140 Ma	Orb-weaver spiders appear.
135 Ma	Rise of the angiosperms. Some of these flowering plants bear structures that attract insects and other animals to spread pollen; other angiosperms are pollinated by wind or water. This innovation causes a major burst of animal coevolution. First freshwater pelomedusid turtles. Earliest krill.
120 Ma	Oldest fossils of heterokonts, including both marine diatoms and silicoflagellates.
115 Ma	First monotreme mammals.
114 Ma	Earliest bees.<ref name="bees-argentina">{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
112 Ma	Xiphactinus, a large predatory fish, appears in the fossil record.
110 Ma	First hesperornithes, toothed diving birds. Earliest limopsid, verticordiid, and thyasirid bivalves.
100 Ma	First ants.<ref>Template:Cite journal</ref>
100–95 Ma	Spinosaurus appears in the fossil record.<ref>Template:Cite journal</ref>
95 Ma	First crocodilians evolve.<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
90 Ma	Extinction of ichthyosaurs. Earliest snakes and nuculanid bivalves. Large diversification in angiosperms: magnoliids, rosids, hamamelidids, monocots, and ginger. Earliest examples of ticks. Probable origins of placental mammals (earliest undisputed fossil evidence is 66 Ma).
86–76 Ma	Diversification of therian mammals.<ref>Template:Cite journal</ref><ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
70 Ma	Multituberculate mammals increase in diversity. First yoldiid bivalves. First possible ungulates (Protungulatum).
68–66 Ma	Tyrannosaurus, the largest terrestrial predator of western North America, appears in the fossil record. First species of Triceratops.<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>

Cenozoic Era

{{#invoke:Labelled list hatnote|labelledList|Main article|Main articles|Main page|Main pages}} Template:More citations needed

File:Patriofelis-mount.jpg

Mount of oxyaenid Patriofelis from the American Museum of Natural History

File:Icaronycteris index.jpg

The bat Icaronycteris appeared 52.2 million years ago

File:Grassflowers.jpg

Grass flowers

File:1064376 - Megafauna - Museu Nacional de História Natural UFRJ - 22 Outubro 2010 - Rio de Janeiro - Brazil.jpg

Reconstructed skeletons of flightless terror bird and ground sloth at the Museu Nacional, Rio de Janeiro

File:Diprotodon optatum (2).jpg

Diprotodon went extinct about 40,000 years ago as part of the Quaternary extinction event, along with every other Australian creature over Template:Cvt.

File:Homo floresiensis v 2-0.jpg

50,000 years ago several different human species coexisted on Earth including modern humans and Homo floresiensis (pictured).

File:PantheraLeoAtrox1 (retouched).jpg

American lions exceeded extant lions in size and ranged over much of North America until 11,000 BP.

Cenozoic era (66 Ma – present)

Date	Event
66 Ma	The Cretaceous–Paleogene extinction event eradicates about half of all animal species, including mosasaurs, pterosaurs, plesiosaurs, ammonites, belemnites, rudist and inoceramid bivalves, most planktic foraminifers, and all of the dinosaurs excluding the birds.<ref>Template:Cite journal</ref>
66 Ma	Rapid dominance of conifers and ginkgos in high latitudes, along with mammals becoming the dominant species. First psammobiid bivalves. Earliest rodents. Rapid diversification in ants.
63 Ma	Evolution of the creodonts, an important group of meat-eating (carnivorous) mammals.
62 Ma	Evolution of the first penguins.
60 Ma	Diversification of large, flightless birds. Earliest true primates,Template:Who along with the first semelid bivalves, edentate, carnivoran and lipotyphlan mammals, and owls. The ancestors of the carnivorous mammals (miacids) were alive.Template:Citation needed
59 Ma	Earliest sailfish appear.
56 Ma	Gastornis, a large flightless bird, appears in the fossil record.
55 Ma	Modern bird groups diversify (first song birds, parrots, loons, swifts, woodpeckers), first whale (Himalayacetus), earliest lagomorphs, armadillos, appearance of sirenian, proboscidean mammals in the fossil record. Flowering plants continue to diversify. The ancestor (according to theory) of the species in the genus Carcharodon, the early mako shark Isurus hastalis, is alive. Ungulates split into artiodactyla and perissodactyla, with some members of the former returning to the sea.
52 Ma	First bats appear (Onychonycteris).
50 Ma	Peak diversity of dinoflagellates and nannofossils, increase in diversity of anomalodesmatan and heteroconch bivalves, brontotheres, tapirs, rhinoceroses, and camels appear in the fossil record, diversification of primates.
40 Ma	Modern-type butterflies and moths appear. Extinction of Gastornis. Basilosaurus, one of the first of the giant whales, appeared in the fossil record.
38 Ma	Earliest bears.
37 Ma	First nimravid ("false saber-toothed cats") carnivores — these species are unrelated to modern-type felines. First alligators and ruminants.
35 Ma	Grasses diversify from among the monocot angiosperms; grasslands begin to expand. Slight increase in diversity of cold-tolerant ostracods and foraminifers, along with major extinctions of gastropods, reptiles, amphibians, and multituberculate mammals. Many modern mammal groups begin to appear: first glyptodonts, ground sloths, canids, peccaries, and the first eagles and hawks. Diversity in toothed and baleen whales.
33 Ma	Evolution of the thylacinid marsupials (Badjcinus).
30 Ma	First balanids and eucalypts, extinction of embrithopod and brontothere mammals, earliest pigs and cats.
28 Ma	Paraceratherium appears in the fossil record, the largest terrestrial mammal that ever lived. First pelicans.
25 Ma	Pelagornis sandersi appears in the fossil record, the largest flying bird that ever lived.
25 Ma	First deer.
24 Ma	First pinnipeds.
23 Ma	Earliest ostriches, trees representative of most major groups of oaks have appeared by now.<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref>
20 Ma	First giraffes, hyenas, and giant anteaters, increase in bird diversity.
17 Ma	First birds of the genus Corvus (crows).
15 Ma	Genus Mammut appears in the fossil record, first bovids and kangaroos, diversity in Australian megafauna.
10 Ma	Grasslands and savannas are established, diversity in insects, especially ants and termites, horses increase in body size and develop high-crowned teeth, major diversification in grassland mammals and snakes.
9.5 Ma {{ safesubst:#invoke:Unsubst	$B= Template:Fix }}	Great American Interchange, where various land and freshwater faunas migrated between North and South America. Armadillos, opossums, hummingbirds Phorusrhacids, Ground Sloths, Glyptodonts, and Meridiungulates traveled to North America, while horses, tapirs, saber-toothed cats, jaguars, bears, coaties, ferrets, otters, skunks and deer entered South America.
9 Ma	First platypuses.
6.5 Ma	First hominins (Sahelanthropus).
6 Ma	Australopithecines diversify (Orrorin, Ardipithecus).
5 Ma	First tree sloths and hippopotami, diversification of grazing herbivores like zebras and elephants, large carnivorous mammals like lions and the genus Canis, burrowing rodents, kangaroos, birds, and small carnivores, vultures increase in size, decrease in the number of perissodactyl mammals. Extinction of nimravid carnivores. First leopard seals.
4.8 Ma	Mammoths appear in the fossil record.
4.5 Ma	Marine iguanas diverge from land iguanas.
4 Ma	Australopithecus evolves. Stupendemys appears in the fossil record as the largest freshwater turtle, first modern elephants, giraffes, zebras, lions, rhinoceros and gazelles appear in the fossil record
3.6 Ma	Blue whales grow to modern size.
3 Ma	Earliest swordfish.
2.7 Ma	Paranthropus evolves.
2.5 Ma	Earliest species of Arctodus and Smilodon evolve.
2 Ma	First members of genus Homo, Homo Habilis, appear in the fossil record. Diversification of conifers in high latitudes. The eventual ancestor of cattle, aurochs (Bos primigenus), evolves in India.
1.7 Ma	Australopithecines go extinct.
1.2 Ma	Evolution of Homo antecessor. The last members of Paranthropus die out.
1.0 Ma	First coyotes.
810 ka	First wolves
600 ka	Evolution of Homo heidelbergensis.
400 ka	First polar bears.
350 ka	Evolution of Neanderthals.
300 ka	Gigantopithecus, a giant relative of the orangutan from Asia dies out.
250 ka	Anatomically modern humans appear in Africa.<ref>Template:Cite journal</ref><ref>Template:Cite press release</ref><ref>Template:Cite journal</ref> Around 50 ka they start colonising the other continents, replacing Neanderthals in Europe and other hominins in Asia.
70 ka	Genetic bottleneck in humans (Toba catastrophe theory).
40 ka	Last giant monitor lizards (Varanus priscus) die out.
35–25 ka	Extinction of Neanderthals. Domestication of dogs.
15 ka	Last woolly rhinoceros (Coelodonta antiquitatis) are believed to have gone extinct.
11 ka	Short-faced bears vanish from North America, with the last giant ground sloths dying out. All Equidae become extinct in North America. Domestication of various ungulates.
10 ka	Holocene epoch starts<ref>{{#invoke:citation/CS1|citation	CitationClass=web }}</ref> after the Last Glacial Maximum. Last mainland species of woolly mammoth (Mammuthus primigenus) die out, as does the last Smilodon species.
8 ka	The giant lemur dies out.

See also

• Evolutionary history of plants (timeline)
• Geologic time scale
• History of Earth
• Sociocultural evolution
• Timeline of human evolution

References

Bibliography

• Template:Cite book
• Template:Cite book
• Template:Cite book
• Template:Cite book
• Template:Cite book
• Template:Cite book
• Template:Cite book

Further reading

• Template:Cite book

External links

• {{#invoke:citation/CS1|citation

|CitationClass=web }}

• {{#invoke:citation/CS1|citation

|CitationClass=web }} Explore complete phylogenetic tree interactively

• {{#invoke:citation/CS1|citation

|CitationClass=web }}

• {{#invoke:citation/CS1|citation

|CitationClass=web }}

• {{#invoke:citation/CS1|citation

|CitationClass=web }} Interactive timeline from Big Bang to present

• {{#invoke:citation/CS1|citation

|CitationClass=web }} Sequence of Plant Evolution

• {{#invoke:citation/CS1|citation

|CitationClass=web }} Sequence of Animal Evolution

• {{#invoke:citation/CS1|citation

|CitationClass=web }}

• Template:Cite AV media
• {{#invoke:citation/CS1|citation

|CitationClass=web }}

• Art of the Nature Timelines on Wikipedia

{{#invoke:Navbox|navbox}} Template:Earth Template:Portal bar