Geranium
Template:Short description Template:About Template:Distinguish Template:Automatic taxobox
Geranium is a genus of 422 species of annual, biennial, and perennial plants that are commonly known as geraniums or cranesbills. They are found throughout the temperate regions of the world and the mountains of the tropics, with the greatest diversity in the eastern part of the Mediterranean region.<ref name="Allaby">Template:Cite book</ref>
The palmately cleft leaves are broadly circular in form. The flowers have five petals and are coloured white, pink, purple, or blue, often with distinctive veining.<ref name="Allaby" /> Geraniums will grow in any soil as long as it is not waterlogged.<ref name="Tibballs">Template:Cite book</ref> Propagation is by semiripe cuttings in summer, by seed, or by division in autumn or spring.<ref>Template:Cite book</ref>
Geraniums are eaten by the larvae of some Lepidoptera species including brown-tail, ghost moth, and mouse moth. At least several species of Geranium are gynodioecious.<ref>Hessing, M.B. 1989. Variation in self-fertility and floral characters of Geranium caespitosum (Geraniaceae) along an elevational gradient. Plant Systematics and Evolution 166:225-241.</ref><ref>Van Etten and Chang 2014. Frequency-dependent pollinator discrimination acts against female plants in the gynodioecious Geranium maculatum Annals of Botany 114:1769–1778</ref><ref>Template:Cite journal</ref> The species Geranium viscosissimum (sticky geranium) is considered to be protocarnivorous.<ref name="Spomer">Template:Cite journal</ref>
NameEdit
The genus name is derived from Ancient Greek {{#invoke:Lang|lang}} (géranos) 'crane'. The English name 'cranesbill' derives from the resemblance of the fruit capsule of some of the species to a crane's head and bill. The ovary portion forms the head and the prolonged stigma creates the appearance of a beak.<ref name=":0">Template:Cite book</ref><ref name="APS">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
DescriptionEdit
The flowers are typically five-petaled and white to purple. The leaves are palmate divided into narrow, pointed segments.<ref name=":0" />
The fruit capsule consists of five cells joined to a column produced from the centre of the flower. The cells form lobes which eventually separate, each containing one seed.<ref name=":0" /> When the fruit is ripe, the beak-like stigma springs open and casts the ovoid, streamlined seeds some distance, dispersing the seeds.<ref>Template:Cite journal</ref>
Confusion with PelargoniumEdit
Confusingly, "geranium" is also the common name of members of the genus Pelargonium, which are also in the family Geraniaceae and are widely grown as horticultural bedding plants. Linnaeus originally included all the species in one genus, Geranium, but they were later separated into two genera by Charles L’Héritier in 1789.<ref>Template:Cite book</ref> Other former members of the genus are now classified in Erodium, including the plants known as filarees in North America.<ref>Template:Cite book</ref>
The term "hardy geranium" is often applied to horticultural Geraniums to distinguish them from the Pelargoniums, which are not winter-hardy in temperate horticulture.<ref name="Parer">Template:Cite book</ref><ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> However, not all Geranium species are winter-hardy (see below).<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
The shape of the flowers offers one way of distinguishing between the two genera Geranium and Pelargonium. Geranium flowers have five very similar petals, and are thus radially symmetrical (actinomorphic), whereas Pelargonium (and also Erodium) flowers have two upper petals which are different from the three lower petals, so the flowers have a single plane of symmetry (zygomorphic).<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref><ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
CultivationEdit
A number of geranium species are cultivated for horticultural use and for pharmaceutical products. Some of the more commonly grown species include: Template:Div col
- Geranium albanum (crested cranesbill)
- Geranium cinereum
- Geranium clarkei (Clarke's geranium)
- Geranium dalmaticum
- Geranium endressii (Endres's cranesbill)
- Geranium erianthum (wooly geranium)
- Geranium fremontii (Fremont's geranium)
- Geranium himalayense, often sold under Geranium grandiflorum
- Geranium ibericum (Caucasus geranium),
- Geranium macrorrhizum (bigroot cranesbill or bigroot geranium)
- Geranium maculatum (wild geranium)
- Geranium maderense (giant herb robert)
- Geranium × magnificum (showy geranium)
- Geranium phaeum (dusky cranesbill)
- Geranium platypetalum (broad-petaled geranium)
- Geranium pratense (meadow cranesbill)
- Geranium psilostemon (Armenian cranesbill)
- Geranium renardii (Renard geranium)
- Geranium sanguineum (bloody cranesbill)
- Geranium subcaulescens (grey cranesbill)
- Geranium sylvaticum (wood cranesbill)
All the above species are perennials and generally winter-hardy plants, grown for their attractive flowers and foliage. They are long-lived and most have a mounding habit, with palmately lobed foliage. Some species have spreading rhizomes. They are normally grown in part shade to full sun, in well-draining but moisture retentive soils, rich in humus.<ref name=Phillips1993>Template:Citation</ref> Other perennial species grown for their flowers and foliage include: Geranium argenteum, G. eriostemon, G. farreri, G. nodosum, G. procurrens, G. pylzowianum, G. renardii, G. traversii, G. tuberosum, G. versicolor, G. wallichianum, and G. wlassovianum. Some of these are not winter-hardy in cold areas and are grown in specialized gardens like rock gardens.<ref name=Jelitto1990>Template:Citation</ref> Geranium 'Johnson's Blue' is a hybrid between G. himalayense (southwestern China), with G. pratense (European meadow cranesbill).<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
CultivarsEdit
The following hybrid cultivars have gained the Royal Horticultural Society's Award of Garden Merit (other cultivars are dealt with under their species name - see above):-<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> Template:Div col
- 'Ann Folkard'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'A. T. Johnson' (G. × oxonianum)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Ballerina' <ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Blue Cloud'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- Blue Sunrise='Blogold' (PBR) <ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Brookside' <ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Danny Boy'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Dilys'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Gypsy' (G. × lindavicum)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Ivan' <ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Mavis Simpson'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Nimbus' <ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Orion'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- Patricia='Brempat'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- Rothbury Gem='Gerfos'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- Rozanne='Gerwat'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Russell Prichard'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Sirak'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Wageningen'<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref>
- 'Wargrave Pink' (G. × oxonianum)<ref>{{#invoke:citation/CS1|citation
|CitationClass=web }}</ref> Template:Div col end
GalleryEdit
- Geranium maculatum - wild cranesbill - desc-flower straight on.jpg
- Wild Geranium (Geranium maculatum).jpg
- Illustration Geranium phaeum0.jpg
Geranium phaeum - from Thomé Flora von Deutschland, Österreich und der Schweiz 1885
- Geranium platypetalum1.jpg
- Geranium sanguineum0.jpg
- Geranium pratense (Meadow Cranesbill).jpg
Geranium pratense (meadow cranesbill)
- Geranium-robertianum(Samen).jpg
Geranium robertianum (herb robert)
- GeraniumMaderense.jpg
- Wildgeranium.jpg
- Starr 980718-1820 Geranium arboreum.jpg
- Płatek bodziszka.jpgGeranium petal under the microscopeFile:Geranios Rosas y Rojos.jpgGeranium
- Geranium (probably 'Johnson's Blue').jpg
Geranium (possibly 'Johnson's Blue')
- גרניון גזור.jpg