Motility
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Motility is the ability of an organism to move independently using metabolic energy. This biological concept encompasses movement at various levels, from whole organisms to cells and subcellular components.
Motility is observed in animals, microorganisms, and even some plant structures, playing crucial roles in activities such as foraging, reproduction, and cellular functions. It is genetically determined but can be influenced by environmental factors.
In multicellular organisms, motility is facilitated by systems like the nervous and musculoskeletal systems, while at the cellular level, it involves mechanisms such as amoeboid movement and flagellar propulsion. These cellular movements can be directed by external stimuli, a phenomenon known as taxis. Examples include chemotaxis (movement along chemical gradients) and phototaxis (movement in response to light).
Motility also includes physiological processes like gastrointestinal movements and peristalsis. Understanding motility is important in biology, medicine, and ecology, as it impacts processes ranging from bacterial behavior to ecosystem dynamics.
DefinitionsEdit
Motility, the ability of an organism to move independently, using metabolic energy,<ref name="rupress">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref><ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref> can be contrasted with sessility, the state of organisms that do not possess a means of self-locomotion and are normally immobile. Motility differs from mobility, the ability of an object to be moved.
The term vagility means a lifeform that can be moved but only passively; sessile organisms including plants and fungi often have vagile parts such as fruits, seeds, or spores which may be dispersed by other agents such as wind, water, or other organisms.<ref name="nerd word">{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
Motility is genetically determined,<ref>Template:Cite book</ref> but may be affected by environmental factors such as toxins. The nervous system and musculoskeletal system provide the majority of mammalian motility.<ref>Template:Cite book</ref><ref>Template:Cite book</ref><ref>Template:Cite journal</ref>
In addition to animal locomotion, most animals are motile, though some are vagile, described as having passive locomotion. Many bacteria and other microorganisms, including even some viruses,<ref>Template:Cite journal</ref> and multicellular organisms are motile; some mechanisms of fluid flow in multicellular organs and tissue are also considered instances of motility, as with gastrointestinal motility. Motile marine animals are commonly called free-swimming,<ref>Template:Cite journal</ref><ref>Template:Cite journal</ref><ref>Template:Cite journal</ref> and motile non-parasitic organisms are called free-living.<ref>{{#invoke:citation/CS1|citation |CitationClass=web }}</ref>
Motility includes an organism's ability to move food through its digestive tract. There are two types of intestinal motility – peristalsis and segmentation.<ref>Template:EMedicine</ref> This motility is brought about by the contraction of smooth muscles in the gastrointestinal tract which mix the luminal contents with various secretions (segmentation) and move contents through the digestive tract from the mouth to the anus (peristalsis).<ref>Template:Cite book</ref>
Cellular levelEdit
At the cellular level, different modes of movement exist:
- amoeboid movement, a crawling-like movement, which also makes swimming possible<ref>Template:Cite journal</ref><ref>Template:Cite journal</ref>
- filopodia, enabling movement of the axonal growth cone<ref name="Gilbert">Template:Cite book</ref>
- flagellar motility, a swimming-like motion (observed for example in spermatozoa, propelled by the regular beat of their flagellum, or the E. coli bacterium, which swims by rotating a helical prokaryotic flagellum)
- gliding motility
- swarming motility
- twitching motility, a form of motility used by bacteria to crawl over surfaces using grappling hook-like filaments called type IV pili.
Many cells are not motile, for example Klebsiella pneumoniae and Shigella, or under specific circumstances such as Yersinia pestis at 37 °C.Template:Citation needed
MovementsEdit
Events perceived as movements can be directed:
- along a chemical gradient (see chemotaxis)
- along a temperature gradient (see thermotaxis)
- along a light gradient (see phototaxis)
- along a magnetic field line (see magnetotaxis)
- along an electric field (see galvanotaxis)
- along the direction of the gravitational force (see gravitaxis)
- along a rigidity gradient (see durotaxis)
- along a gradient of cell adhesion sites (see haptotaxis)
- along other cells or biopolymers
- Muscular system.svg
Muscles give the ability for voluntary movement, and involuntary movement as in muscle spasms and reflexes. At the level of the muscular system, motility is a synonym for locomotion.<ref>Template:Cite book</ref><ref>Template:Cite book</ref>
- Cheetah chase.jpg
The record speeds cheetahs hold are owed in large to their muscle motility.
- Phototropism.jpg
The shoots of plants move by growing towards light. This is known as positive phototropism. The roots grow away from light. This is known as negative phototropism.
- FAGOCITOSI BY RAFF.gif
Monocytes and macrophages of the immune system engulf Bacteria by extending their pseudopodia. Note that this cartoon is not an accurate representation of phagocytosis.
- Protein translation.gif
Motility at the sub-cellular level. This depicts translation - a motile nanoscale molecular process using protein dynamics.