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Hellbender
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{{Short description|Species of amphibian}} {{About|amphibians|other uses}} {{Speciesbox | name = Hellbender | fossil_range = <br /> {{longitem|style=line-height:1.25em|{{nowrap|[[Irvingtonian|Middle Pleistocene]] – [[Holocene|present]]}} {{nowrap|(~625,000–0 [[Before Present|YBP]])<ref name=Bredehoeft&Schubert2015>{{cite journal |last1=Bredehoeft |first1=Keila E. |last2=Schubert |first2=Blaine W. |year=2015 |title=A re-evaluation of the Pleistocene hellbender, ''Cryptobranchus guildayi ''|journal=[[Journal of Herpetology]] |volume=49 |pages=157–160 |s2cid=84731832 |doi=10.1670/12-222 }}</ref>}}}} | status = VU | status_system = IUCN3.1 | status_ref = <ref name=iucn>{{cite iucn |author=IUCN SSC Amphibian Specialist Group. |year=2022 |title=''Cryptobranchus alleganiensis'' |page=e.T59077A82473431 |doi=10.2305/IUCN.UK.2022-2.RLTS.T59077A82473431.en}}</ref> | status2 = CITES_A3 | status2_system = CITES | status2_ref = | image = Hellbender.jpg | genus = Cryptobranchus | parent_authority = [[Friedrich Sigismund Leuckart|Leuckart]], 1821 | species = alleganiensis | authority = ([[François Marie Daudin|Daudin]], 1803) | subdivision_ranks = Subspecies | subdivision = *'''''C. a. alleganiensis''''' {{small|(Daudin, 1803)}}, eastern hellbender *'''''C. a. bishopi''''' {{small|[[Arnold B. Grobman|Grobman]], 1943}}, Ozark hellbender | range_map = Hellbender.png | range_map_caption = {{legend0|#af3635| Range of eastern hellbender}}<br />(Ozark hellbender not shown) | synonyms = {{Collapsible list|{{specieslist |Salamandra alleganiensis |Daudin, 1803 |Salamandra horrida |[[Benjamin Smith Barton|Barton]], 1808 |Salamandra gigantea |Barton, 1808 |Salamandra maxima |Barton, 1808 |Molge gigantea |[[Blasius Merrem|Merrem]], 1820 |Cryptobranchus salamandroides |[[Friedrich Sigismund Leuckart|Leuckart]], 1821 |Urotropis mucronata |[[Constantine Samuel Rafinesque|Rafinesque]], 1822 |Abranchus alleghaniensis |[[Richard Harlan|Harlan]], 1825 |Protonopsis horrida |[[Daniel H. Barnes|Barnes]], 1826 |Salamandrops gigantea |[[Johann Georg Wagler|Wagler]], 1830 |Eurycea mucronata |Rafinesque, 1832 |Menopoma fuscum |[[John Edwards Holbrook|Holbrook]], 1842 |Cryptobranchus alleghaniensis |[[Edward Drinker Cope|Cope]], 1887 |Cryptobranchus alleganiensis |[[Leonhard Stejneger|Stejneger]] & [[Thomas Barbour|Barbour]], 1917 |Cryptobranchus terrasodactylus |[[Vera Wellborn|Wellborn]], 1936 |Cryptobranchus bishopi |[[Arnold B. Grobman|Grobman]], 1943 |Cryptobranchus guildayi |[[J. Alan Holman|Holman]], 1977 }} }} | synonyms_ref = <ref>{{cite report |last=Dundee |first=Harold A. |year=1971 |section=Cryptobranchus alleganiensis |title=Catalogue of American Amphibians and Reptiles |publisher=[[American Society of Ichthyologists and Herpetologists]] |id=Account 101 }}</ref><ref name=Nick/><ref>{{cite book |author1-link=Leonhard Stejneger |last1=Stejneger |first1=L. |author2-link=Thomas Barbour |last2=Barbour |first2=T. |year=1917 |title=A Check List of North American Amphibians and Reptiles |place=Cambridge, MA |publisher=Harvard University Press |section=Cryptobranchus alleganiensis |page=7 }}</ref> }} The '''hellbender''' ('''''Cryptobranchus alleganiensis'''''), also known as the '''hellbender salamander''', is a [[species]] of aquatic [[giant salamander]] [[Endemism|endemic]] to the eastern and central United States. It is the largest salamander in [[North America]]. A member of the [[Family (biology)|family]] [[Cryptobranchidae]], the hellbender is the only [[extant taxon|extant]] member of the genus ''Cryptobranchus''. Other closely related salamanders in the same family are in the genus ''[[Andrias]]'', which contains the [[Japanese giant salamander|Japanese]] and [[Chinese giant salamander]]s. The hellbender is much larger than any other salamander in its geographic range, and employs an unusual adaption for respiration through [[cutaneous gas exchange]] via [[capillaries]] found in its [[Anatomical terms of location#Medial and lateral|lateral]] skin folds. It fills a particular [[Niche (ecology)|niche]]—both as a [[predator]] and [[prey]]—in its [[ecosystem]], which either it or its ancestors have occupied for around 65 million years. The species is listed as [[Vulnerable species|Vulnerable]] on the [[IUCN Red List of Threatened Species]] due to the impacts of disease and widespread habitat loss and degradation throughout much of its range.<ref name=iucn/>
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