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Elasmobranchii
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==Description== Members of the Elasmobranchii subclass have no [[swim bladders]], five to seven pairs of [[gill]] clefts opening individually to the exterior, rigid [[dorsal fin]]s, and small [[placoid scale]]s. The teeth are in several series; the upper jaw is not fused to the cranium, and the lower jaw is articulated with the upper. Extant elasmobranchs exhibit several archetypal jaw suspensions: amphistyly, orbitostyly, hyostyly, and euhyostyly. In amphistyly, the palatoquadrate has a postorbital articulation with the chondrocranium from which ligaments primarily suspend it anteriorly. The hyoid articulates with the mandibular arch posteriorly, but it appears to provide little support to the upper and lower jaws. In orbitostyly, the orbital process hinges with the orbital wall and the hyoid provides the majority of suspensory support. In contrast, hyostyly involves an ethmoid articulation between the upper jaw and the cranium, while the hyoid most likely provides vastly more jaw support compared to the anterior ligaments. Finally, in euhyostyly, also known as true hyostyly, the mandibular cartilages lack a ligamentous connection to the cranium. Instead, the hyomandibular cartilages provide the only means of jaw support, while the ceratohyal and basihyal elements articulate with the lower jaw, but are disconnected from the rest of the hyoid.<ref>{{cite journal |doi=10.1002/jmor.10342 |pmid=15880740 |title=Morphology and evolution of the jaw suspension in lamniform sharks |journal=Journal of Morphology |volume=265 |issue=1 |pages=102β19 |year=2005 |last1=Wilga |first1=C.D. |s2cid=45227734 }}</ref><ref>{{cite journal |doi=10.1093/icb/icm029 |pmid=21672820 |title=Evolution and ecology of feeding in elasmobranchs |journal=Integrative and Comparative Biology |volume=47 |issue=1 |pages=55β69 |year=2007 |last1=Wilga |first1=C. D. |last2=Motta |first2=P. J. |last3=Sanford |first3=C. P. |doi-access=free }}</ref><ref>{{cite journal |last1=Wilga |first1=Cheryl A.D. |year=2008 |title=Evolutionary divergence in the feeding mechanism of fishes |journal=Acta Geologica Polonica |volume=58 |issue=2 |pages=113β20 |url=https://geojournals.pgi.gov.pl/agp/article/view/9981 |access-date=2017-05-24 |archive-url=https://web.archive.org/web/20180819031736/https://geojournals.pgi.gov.pl/agp/article/view/9981 |archive-date=2018-08-19 |url-status=live }}</ref> The eyes have a [[tapetum lucidum]]. The inner margin of each pelvic fin in the male fish is grooved to constitute a [[clasper]] for the transmission of [[sperm]]. These fish are widely distributed in [[tropical]] and [[temperate]] waters.<ref>{{cite book | last = Bigelow| first = Henry B. | author-link = Henry Bryant Bigelow|author2=Schroeder, William C.| title = Fishes of the Western North Atlantic | publisher = Sears Foundation for Marine Research, Yale University | year = 1948 | pages = 64β65 | asin = B000J0D9X6}}</ref> Many fish maintain buoyancy with [[swim bladder]]s. However elasmobranchs lack swim bladders, and maintain buoyancy instead with large livers that are full of oil.<ref>Oguri, M (1990) [http://spo.nwr.noaa.gov/tr90opt.pdf "A review of selected physiological characteristics unique to elasmobranchs"] {{Webarchive|url=https://web.archive.org/web/20130218135249/http://spo.nwr.noaa.gov/tr90opt.pdf |date=2013-02-18 }} In: ''Elasmobranchs as living resources: advances in the biology, ecology, systematics and the status of the fisheries'', eds. J. H. L. Pratt, S. H. Gruber and T. Taniuchi, US Department of Commerce, NOAA technical report NMFS 90, pp.49β54.</ref> This stored oil may also function as a nutrient when food is scarce.<ref name=Hoenig1990>Hoenig, J.M. and Gruber, S.H. (1990) [http://spo.nwr.noaa.gov/tr90opt.pdf "Life-history patterns in the elasmobranchs: implications for fisheries management"] {{Webarchive|url=https://web.archive.org/web/20130218135249/http://spo.nwr.noaa.gov/tr90opt.pdf |date=2013-02-18 }} In: ''Elasmobranchs as living resources: advances in the biology, ecology, systematics and the status of the fisheries'', eds. J. H. L. Pratt, S. H. Gruber and T. Taniuchi, US Department of Commerce, NOAA technical report NMFS 90, pp.1β16.</ref><ref>{{cite journal |doi=10.1017/S0025315400038017 |title=The density of elasmobranchs |journal=Journal of the Marine Biological Association of the United Kingdom |volume=49 |issue=4 |pages=913 |year=2009 |last1=Bone |first1=Q. |last2=Roberts |first2=B. L. |s2cid=85871565 }}</ref>
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