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Local extinction
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==Discussion== {{see also|Biogeography|Metapopulation|Patch dynamics|Habitat fragmentation}} [[Glacial period|Glaciation]] is one factor that leads to local extinction. This was the case during the [[Quaternary glaciation|Pleistocene glaciation]] event in North America. During this period, most of the native North American species of earthworm were killed in places covered by glaciation. This left them open for colonization by European earthworms brought over in soil from Europe.<ref>{{Cite journal|last=Gates|first=G. E.|date=May 1949|title=Miscellanea Megadrilogica|url=http://dx.doi.org/10.1086/281596|journal=The American Naturalist|volume=83|issue=810|pages=139β152|doi=10.1086/281596|s2cid=222328616|issn=0003-0147|url-access=subscription}}</ref> Species naturally become extinct from islands over time; this can be either local extinction if the species also occurs elsewhere, or in cases of [[endemism|island endemism]], outright extinction. The number of species an island can support is limited by its geographical size. Because many islands were relatively recently formed due to climate change at the end of the [[Pleistocene]] when the sea level rose, and these islands most likely had the same complement of species as found on the mainland, counting the species which still survive at present on a statistically large enough amount of islands will give the parameters with which certain groups of species such as plants or birds will become less biodiverse on a given island over a given period of time, depending on its size. The same calculations can also be applied to determine when species will disappear from nature parks ('islands' in many senses), mountain tops and mesas (see [[sky island]]s), forest remnants or other such distributional patches. This research also demonstrates that certain species are more prone to extinction than others, a species has an intrinsic extinction-ability (incidence function).<ref>{{cite journal |last1=Diamond |first1=Jared M. |author-link1=Jared Diamond |date=November 1972 |title=Biogeographic Kinetics: Estimation of Relaxation Times for Avifaunas of Southwest Pacific Islands |url=http://jareddiamond.org/Jared_Diamond/Further_Reading_files/Diamond%201972_1.pdf |journal=[[Proceedings of the National Academy of Sciences of the USA]] |volume=69 |issue=11 |pages=3199β3203 |doi=10.1073/pnas.69.11.3199 |pmid=16592024 |pmc=389735 |bibcode=1972PNAS...69.3199D |access-date=21 November 2021|doi-access=free }}</ref><ref>{{cite journal |last1=Diamond |first1=Jared M. |author-link1=Jared Diamond |date=1975 |title=The Island Dilemma: Lessons of Modern Biogeographic Srudies for the Design of Natural Reserves |url=http://jareddiamond.org/Jared_Diamond/Further_Reading_files/Diamond%201975.pdf |journal=Biological Conservation |volume=7 |issue=2 |pages=129β146 |doi=10.1016/0006-3207(75)90052-X |bibcode=1975BCons...7..129D |access-date=21 November 2021}}</ref> Some species exploit or require transient or [[disturbance (ecology)|disturbed]] habitats, such as vernal pools, a human gut, or burnt woodland after [[forest fire]]s, and are characterised by highly fluctuating population numbers and shifting distributional patterns. Many natural ecosystems cycle through a standard [[succession (ecology)|succession]], [[pioneer species]] disappear from a region as the ecosystem matures and reaches a [[climax community]]. A local extinction can be useful for research: in the case of the [[bay checkerspot butterfly]], scientists, including [[Paul R. Ehrlich]], chose not to intervene as a population disappeared from an area in order to study the process.<ref name="Holsinger">Holsinger, Kent. "[http://darwin.eeb.uconn.edu/eeb310/lecture-notes/checkerspot/node4.html Local extinction] {{webarchive|url=https://web.archive.org/web/20070311042359/http://darwin.eeb.uconn.edu/eeb310/lecture-notes/checkerspot/node4.html |date=2007-03-11 }}". ''Population Viability Analysis: Bay Checkerspot Butterfly''. URL accessed August 11, 2006.</ref> Many crocodilian species have experienced localized extinction, particularly the [[saltwater crocodile]] (''Crocodylus porosus''), which has been extirpated from Vietnam, Thailand, Java, and many other areas.<ref>{{Cite book|last1=Webb|first1=Grahame|url=https://d1wqtxts1xzle7.cloudfront.net/53234109/croc_2.pdf?1495471558=&response-content-disposition=inline%3B+filename%3DCroc.pdf&Expires=1617324076&Signature=RWpZNgowvcRiT5~eVvXnNiIu4T-dw9tkY6F5~IspzSAVjJ6E1BdwdQwbRi9ukFVPINwW8-SlFHgjrkICXZmo7PSqDHMdirdfTcVNmVSS2HtCLP7vE3ys3wvSiQlMPCvG0q~kvrEm9gY9V6BMD0mN41ek8JnlWKEk6yRdvsgthCCzW~886ei1uZngRClj33oltALQCo4rfymrxippXRT8Hhomz6JLz-cpYdA~1~w-lz~VcmVjMkArvumOvQ6K3yd6PK-h2nvcU5HFYniQGEMtR-XPLzFcPhOb5VE9kl01Wd2wYhTt3DmO0nrmHAyoQVLb86Jo9LpR8sDD0utKUCWjlw__&Key-Pair-Id=APKAJLOHF5GGSLRBV4ZA|title=Saltwater Crocodile ''Crocodylus porosus''|last2=Manolis|first2=S|last3=Brien|first3=Matthew|publisher=Crocodiles|year=2010|pages=99β113}}{{dead link|date=May 2021|bot=medic}}{{cbignore|bot=medic}}</ref> [[File:Wolf Canis lupus Wildpark Poing-09.jpg|thumb|''Canis lupus'']] Major environmental events, such as volcanic eruptions, may lead to large numbers of local extinctions, such as with the [[1980 Mount St. Helens eruption]], which led to a [[fern spike]] extinction. [[File:Durvillaea potatorum Bull kelp P2153881.JPG|thumb|Bull Kelp]] Heat waves can lead to local extinction. In New Zealand, during the summer of 2017β2018, [[sea surface temperature]]s around parts of [[South Island]] exceeded {{convert|23|C|abbr=on}}, which was well above normal. Air temperatures were also high, exceeding {{convert|30|C|abbr=on}}. These high temperatures, coupled with small wave height, led to the local extinction of [[Durvillaea|bull kelp (''Durvillaea'' spp.)]] from [[Lyttelton Harbour#Bays and headlands|Pile Bay]].<ref name=":0">{{Cite journal |last1=Thomsen|first1=Mads S. |last2=Mondardini |first2=Luca |last3=Alestra |first3=Tommaso |last4=Gerrity |first4=Shawn |last5=Tait |first5=Leigh |last6=South |first6=Paul M. |last7=Lilley |first7=Stacie A. |last8=Schiel |first8=David R. |name-list-style=amp |date=March 2019 |title=Local Extinction of Bull Kelp (''Durvillaea'' spp.) Due to a Marine Heatwave |journal=Frontiers in Marine Science |volume=6 |doi=10.3389/fmars.2019.00084 |issn=2296-7745 |doi-access=free}}</ref> Lagoa Santa, a lake located in [[Lagoa Santa, Minas Gerais|Lagoa Santa, Brazil]], has lost almost 70% of the local fish species over the last 150 years. These include ''[[Acestrorhynchus lacustris]]'', ''[[Astyanax fasciatus]]'', and ''[[Characidium zebra]]''. This could be caused by the introduction of non-native species, like ''[[Redbreast tilapia|Tilapia rendalli]]'', into the lagoon, changes in water level and organic pollution.<ref>{{Cite journal|last1=Pompeu|first1=Paulo dos Santos|last2=Alves|first2=Carlos Bernardo Mascarenhas|date=December 2003|title=Local fish extinction in a small tropical lake in Brazil|journal=Neotropical Ichthyology|language=en|volume=1|issue=2|pages=133β135|doi=10.1590/S1679-62252003000200008|issn=1679-6225|doi-access=free}}</ref> Local extinctions can be reversed, in some cases artificially. Wolves are a species that have been [[Reintroduction|reintroduced]] into parts of their historical range. This has happened with red wolves (''[[Red wolf|Canis rufus]]'') in the United States in the late 1980s and also grey wolves in Yellowstone National Park in the mid-1990s. There have been talks of reintroducing wolves in Scotland, Japan, and Mexico.<ref>{{Cite journal|last1=Arts|first1=Koen|last2=Fischer|first2=Anke|last3=Wal|first3=RenΓ© van der|date=2016|title=Boundaries of the wolf and the wild: a conceptual examination of the relationship between rewilding and animal reintroduction|url=https://onlinelibrary.wiley.com/doi/abs/10.1111/rec.12309|journal=Restoration Ecology|language=en|volume=24|issue=1|pages=27β34|doi=10.1111/rec.12309|bibcode=2016ResEc..24...27A |s2cid=86338008|issn=1526-100X|url-access=subscription}}</ref>
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