Editing Muscular hydrostat (section)

==Muscular anatomy==
[[File:Elephas Maximus Trunk Closeup.jpg|thumb|upright 0.6|Closeup of the trunk of an [[Asian elephant]]]]
Muscles provide the force to move a muscular hydrostat.  Since muscles are only able to produce force by contracting and becoming shorter, different groups of muscles have to work against each other, with one group relaxing and lengthening as the other group provides the force by contracting.  Such complementary muscle groups are termed [[antagonistic pairs]].

The muscle fibers in a muscular hydrostat are oriented in three directions: parallel to the long axis, perpendicular to the long axis, and wrapped obliquely around the long axis.<ref name=Kier&Smith1985>{{Cite journal | last1 = Kier | first1 = W. M. | last2 = Smith | first2 = K. K. | doi = 10.1111/j.1096-3642.1985.tb01178.x | title = Tongues, tentacles and trunks: The biomechanics of movement in muscular-hydrostats | journal = Zoological Journal of the Linnean Society | volume = 83 | issue = 4 | pages = 307–324| year = 1985 }}</ref><ref name=Smith&Kier1989>{{cite journal|last=Smith|first=Kathleen K.|author2=William M. Kier|title=Trunks, tongues, and tentacles: Moving with skeletons of muscle|journal=American Scientist|date=Jan–Feb 1989|volume=77|issue=1|pages=28–35}}</ref>

The muscles parallel to the long axis are arranged in longitudinal bundles.  The more peripherally these are located, the more elaborate bending movements are possible.  A more peripheral distribution is found in [[tetrapod]] tongues, [[octopus]] arms, [[nautilus]] tentacles, and [[Elephant#Trunk|elephant trunks]].  Tongues that are adapted for protrusion typically have centrally located longitudinal fibers.  These are found in [[snake]] tongues, many [[lizard]] tongues, and the mammalian [[anteater]]s.

The muscles perpendicular to the long axis may be arranged in a transverse, circular, or radial pattern. A transverse arrangement involves sheets of muscle fibers running perpendicular to the long axis, usually alternating between horizontal and vertical orientations. This arrangement is found in the arms and tentacles of squid, octopuses, and in most mammalian tongues.  A radial arrangement involves fibers radiating out in all directions from the center of the organ.  This is found in the tentacles of the [[chambered nautilus]] and in the elephant [[proboscis]] (trunk). A circular arrangement has rings of contractive fibers around the long axis. This is found in many mammalian and lizard tongues along with squid tentacles.

Helical or oblique fibers around the long axis are generally present in two layers with opposite [[chirality]] and wrap around the central core of musculature.