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Ploceidae
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==Taxonomy and systematics== The family Ploceidae was introduced (as Ploceïdes) by Swedish zoologist [[Carl Jakob Sundevall]] in 1836.<ref>{{cite periodical| last=Bock | first=Walter J. | year=1994 | title=History and Nomenclature of Avian Family-Group Names |periodical=Bulletin of the American Museum of Natural History |number=222 | publisher=American Museum of Natural History |location=New York | pages=157, 260 | hdl=2246/830 | url=http://digitallibrary.amnh.org/handle/2246/830 }}<!--Linked page allows download of the 48MB pdf--></ref><ref>{{ cite journal | last=Sundevall | first=Carl Jakob | year=1836 | title=Ornithologiskt system | journal=Kongliga Svenska Vetenskapsakademiens Handlingar | volume=23 | pages=43–130 [74] | url=https://www.biodiversitylibrary.org/page/34523886 }}</ref> Phylogenetic studies have shown that the family is [[sister taxon|sister]] to a clade containing the families [[Viduidae]] and [[Estrildidae]]<ref>{{Cite journal |vauthors=De Silva TN, Peterson AT, Perktas U |date=1 July 2019 |title=An extensive molecular phylogeny of weaverbirds (Aves: Ploceidae) unveils broad nonmonophyly of traditional genera and new relationships |journal=The Auk |volume=136 |issue=3 |doi=10.1093/auk/ukz041 |doi-access=}}</ref> Their common ancestor lived in the middle [[Miocene]] around 18 million years ago.<ref>{{ cite journal | last1=Oliveros | first1=C.H. | display-authors=etal | year=2019 | title=Earth history and the passerine superradiation | journal=Proceedings of the National Academy of Sciences of the United States | volume=116 | issue=16 | pages=7916–7925 | doi=10.1073/pnas.1813206116 | pmid=30936315 | pmc=6475423 | doi-access=free | bibcode=2019PNAS..116.7916O }}</ref> A 2017 [[molecular phylogenetic]] study by Thilina de Silva and collaborators, as well as an expanded study by the same group published in 2019 have indicated that the genus ''[[Ploceus]]'' as currently defined is [[polyphyletic]].<ref name=silva2017>{{Cite journal | last1=De Silva | first1=T.N. | last2=Peterson | first2=A.T. | last3=Bates | first3=J.M. | last4=Fernando | first4=S.W. | last5=Girard | first5=M.G. | date=2017 | title=Phylogenetic relationships of weaverbirds (Aves: Ploceidae): A first robust phylogeny based on mitochondrial and nuclear markers | journal=Molecular Phylogenetics and Evolution | volume=109 | pages=21–32 | doi=10.1016/j.ympev.2016.12.013 | pmid=28012957 | bibcode=2017MolPE.109...21D | s2cid=205841906 | url=https://www.researchgate.net/publication/311908775 }}</ref><ref name=silva2019>{{Cite journal | last1=De Silva | first1=T.N. | last2=Peterson | first2=A.T. | last3=Perktas | first3=U. | date=2019 | title=An extensive molecular phylogeny of weaverbirds (Aves: Ploceidae) unveils broad nonmonophyly of traditional genera and new relationships | journal=The Auk | volume=136 | issue=ukz041 | doi=10.1093/auk/ukz041 | doi-access= }}</ref> A cladogram based on these results is shown below.<ref name=silva2019/> {{clade| style=font-size:90%;line-height:90% |label1='''Ploceidae''' |1={{Clade |1={{Clade |1=''[[Amblyospiza]]'' – thick-billed weaver |2={{clade |1={{clade |1=''[[Sporopipes]]'' – 2 species (weavers) |2={{clade |1=''[[Plocepasser]]'' – 4 species (sparrow-weavers) |2={{clade |1=''[[Philetairus]]'' – sociable weaver |2=''[[Pseudonigrita]]'' – 2 species (social weavers) }} }} }} |2={{clade |1={{clade |1=''[[Dinemellia]]'' – white-headed buffalo weaver |2=''[[Bubalornis]]'' – 2 species (buffalo weavers) }} |2={{clade |1={{clade |1=''[[Euplectes]]'' – 18 species (bishops and widowbirds) |2={{clade |1=''[[Ploceus]]'' – 5 species (Asian weavers) |2={{clade |1=''[[Quelea]]'' – 3 species (queleas) |2={{clade |1=''[[Pachyphantes]]'' – compact weaver |2=''[[Foudia]]'' – 8 species (fodies) }} }} }} }} |2={{clade |1=''[[Ploceus]]'' – 2 species (Sakalava weaver and Nelicourvi weaver) |2=''[[Ploceus]]''+''[[Malimbus]]''+''[[Anaplectes]]'' – 60 + 10 + 2 = 72 species }} }} }} }} }} }} }} ===Genera=== The family includes 15 genera with a total of 122 species.<ref name=ioc>{{cite web| editor1-last=Gill | editor1-first=Frank | editor1-link=Frank Gill (ornithologist) | editor2-last=Donsker | editor2-first=David | editor3-last=Rasmussen | editor3-first=Pamela | editor3-link=Pamela C. Rasmussen | date=December 2023 | title=Old World sparrows, snowfinches, weavers | work=IOC World Bird List Version 14.1 | url=https://www.worldbirdnames.org/bow/weavers/ | publisher=International Ornithologists' Union | access-date=9 February 2024 }}</ref> For more detail, see [[list of Ploceidae species]]. {| class="wikitable" |- ! Image ! Genus !Species |- |[[File:Red-billed Buffalo Weaver.jpg|175px]] |''[[Bubalornis]]'' {{small|A. Smith, 1836}} | * [[Red-billed buffalo weaver]] (''Bubalornis niger'') * [[White-billed buffalo weaver]] (''Bubalornis albirostris'') |- |[[File:Dinemellia dinemelli.jpg|175px]] |''Dinemellia'' {{small|Reichenbach, 1863}} | * [[White-headed buffalo weaver]] (''Dinemellia dinemelli'') |- |[[File:Plocepasser mahali -Baringo Lake, Kenya -male-8.jpg|175px]] |''[[Plocepasser]]'' {{small|A. Smith, 1836}} | * [[Donaldson Smith's sparrow-weaver]] (''Plocepasser donaldsoni'') * [[Chestnut-backed sparrow-weaver]] (''Plocepasser rufoscapulatus'') * [[Chestnut-crowned sparrow-weaver]] (''Plocepasser superciliosus'') * [[White-browed sparrow-weaver]] (''Plocepasser mahali'') |- |[[File:Weaver bird.jpg|175px]] |''Histurgops'' {{small|Reichenow, 1887}} | * [[Rufous-tailed weaver]] (''Histurgops ruficauda'') |- |[[File:Black-capped Social-Weaver - Samburu - Kenya S4E5139 (22836895922).jpg|175px]] |''[[Pseudonigrita]]'' {{small|Reichenow, 1903}} | * [[Grey-capped social weaver]] (''Pseudonigrita arnaudi'') * [[Black-capped social weaver]] (''Pseudonigrita cabanisi'') |- |[[File:Sociable weaver (Philetairus socius).jpg|175px]] |''Philetairus'' {{small|A. Smith, 1837}} | * [[Sociable weaver]] (''Philetairus socius'') |- |[[File:Speckle-fronted Weaver RWD4.jpg|175px]] |''[[Sporopipes]]'' {{small|Cabanis, 1847}} | * [[Speckle-fronted weaver]] (''Sporopipes frontalis'') * [[Scaly-feathered weaver]] (''Sporopipes squamifrons'') |- |[[File:Amblyospiza albifrons, w, vreet netel-dopvrugte, a, Skeerpoort.jpg|175px]] |''Amblyospiza'' {{small|Sundevall, 1850}} | * [[Thick-billed weaver]] (''Amblyospiza albifrons'') |- |[[File:Black-headed weaver (Ploceus cucullatus bohndorffi) male.jpg|175px]] |''[[Ploceus]]'' {{small|Cuvier, 1816}} | * [[Baglafecht weaver]] (''Ploceus baglafecht'') * [[Bannerman's weaver]] (''Ploceus bannermani'') * [[Bates's weaver]] (''Ploceus batesi'') * [[Black-chinned weaver]] (''Ploceus nigrimentus'') * [[Bertram's weaver]] (''Ploceus bertrandi'') * [[Slender-billed weaver]] (''Ploceus pelzelni'') * [[Loango weaver]] (''Ploceus subpersonatus'') * [[Little weaver]] (''Ploceus luteolus'') * [[Spectacled weaver]] (''Ploceus ocularis'') * [[Black-necked weaver]] (''Ploceus nigricollis'') * [[Olive-naped weaver]] (''Ploceus brachypterus'') * [[Strange weaver]] (''Ploceus alienus'') * [[Black-billed weaver]] (''Ploceus melanogaster'') * [[Cape weaver]] (''Ploceus capensis'') * [[Bocage's weaver]] (''Ploceus temporalis'') * [[Eastern golden weaver]] (''Ploceus subaureus'') * [[Holub's golden weaver]] (''Ploceus xanthops'') * [[Orange weaver]] (''Ploceus aurantius'') * [[Heuglin's masked weaver]] (''Ploceus heuglini'') * [[Golden palm weaver]] (''Ploceus bojeri'') * [[Taveta weaver]] (''Ploceus castaneiceps'') * [[Príncipe weaver]] (''Ploceus princeps'') * [[Northern brown-throated weaver]] (''Ploceus castanops'') * [[Southern brown-throated weaver]] (''Ploceus xanthopterus'') * [[Ruvu weaver]] (''Ploceus holoxanthus'') * [[Kilombero weaver]] (''Ploceus burnieri'') * [[Rüppell's weaver]] (''Ploceus galbula'') * [[Northern masked weaver]] (''Ploceus taeniopterus'') * [[Lesser masked weaver]] (''Ploceus intermedius'') * [[Southern masked weaver]] (''Ploceus velatus'') * [[Katanga masked weaver]] (''Ploceus katangae'') * [[Lufira masked weaver]] (''Ploceus ruweti'') * [[Tanzanian masked weaver]] (''Ploceus reichardi'') * [[Vitelline masked weaver]] (''Ploceus vitellinus'') * [[Speke's weaver]] (''Ploceus spekei'') * [[Fox's weaver]] (''Ploceus spekeoides'') * [[Village weaver]] (''Ploceus cucullatus'') * [[Giant weaver]] (''Ploceus grandis'') * [[Vieillot's black weaver]] (''Ploceus nigerrimus'') * [[Chestnut-and-black weaver]] (''Ploceus castaneofuscus'') * [[Weyns's weaver]] (''Ploceus weynsi'') * [[Clarke's weaver]] (''Ploceus golandi'') * [[Juba weaver]] (''Ploceus dichrocephalus'') * [[Black-headed weaver]] (''Ploceus melanocephalus'') * [[Golden-backed weaver]] (''Ploceus jacksoni'') * [[Cinnamon weaver]] (''Ploceus badius'') * [[Chestnut weaver]] (''Ploceus rubiginosus'') * [[Golden-naped weaver]] (''Ploceus aureonucha'') * [[Yellow-mantled weaver]] (''Ploceus tricolor'') * [[Maxwell's black weaver]] (''Ploceus albinucha'') * [[Nelicourvi weaver]] (''Ploceus nelicourvi'') * [[Sakalava weaver]] (''Ploceus sakalava'') * [[Asian golden weaver]] (''Ploceus hypoxanthus'') * [[Compact weaver]] (''Ploceus superciliosus'') * [[Black-breasted weaver]] (''Ploceus benghalensis'') * [[Streaked weaver]] (''Ploceus manyar'') * [[Baya weaver]] (''Ploceus philippinus'') * [[Finn's weaver]] (''Ploceus megarhynchus'') * [[Dark-backed weaver]] (''Ploceus bicolor'') * [[Preuss's weaver]] (''Ploceus preussi'') * [[Yellow-capped weaver]] (''Ploceus dorsomaculatus'') * [[Olive-headed weaver]] (''Ploceus olivaceiceps'') * [[Usambara weaver]] (''Ploceus nicolli'') * [[Brown-capped weaver]] (''Ploceus insignis'') * [[Bar-winged weaver]] (''Ploceus angolensis'') * [[São Tomé weaver]] (''Ploceus sanctithomae'') * [[Yellow-legged weaver]] (''Ploceus flavipes'') |- |[[File:Crested Malimbe - Kakum - Ghana S4E1412 (22229307983).jpg|175px]] |''[[Malimbus]]'' {{small|Vieillot, 1805}} | * [[Red-crowned malimbe]] (''Malimbus coronatus'') * [[Cassin's malimbe]] (''Malimbus cassini'') * [[Rachel's malimbe]] (''Malimbus racheliae'') * [[Gola malimbe]] (''Malimbus ballmanni'') * [[Red-vented malimbe]] (''Malimbus scutatus'') * [[Ibadan malimbe]] (''Malimbus ibadanensis'') * [[Blue-billed malimbe]] (''Malimbus nitens'') * [[Red-headed malimbe]] (''Malimbus rubricollis'') * [[Red-bellied malimbe]] (''Malimbus erythrogaster'') * [[Crested malimbe]] (''Malimbus malimbicus'') |- |[[File:Quelea erythrops -South Africa -building nest-8.jpg|175px]] |''[[Quelea]]'' {{small|Reichenbach, 1850}} | *[[Cardinal quelea]] (''Quelea cardinalis'') *[[Red-headed quelea]] (''Quelea erythrops'') *[[Red-billed quelea]] (''Quelea quelea'') |- |[[File:Red-headed Weaver male RWD.jpg|175px]] |''[[Anaplectes]]'' {{small|Reichenbach, 1863}} | * [[Red-headed weaver]] (''Anaplectes rubriceps'') * [[Red weaver]] (''Anaplectes jubaensis'') |- |[[File:Madagascar fody (Foudia madagascariensis).jpg|175px]] |''[[Foudia]]'' {{small|Reichenbach, 1850}} | *[[Red fody]] or Madagascar fody (''Foudia madagascariensis'') *[[Comoros fody]] or red-headed fody (''Foudia eminentissima'') *[[Aldabra fody]] (''Foudia aldabrana'') *[[Forest fody]] (''Foudia omissa'') *[[Mauritius fody]] (''Foudia rubra'') *[[Seychelles fody]] (''Foudia sechellarum'') *[[Rodrigues fody]] (''Foudia flavicans'') *[[Réunion fody]] (''Foudia delloni'') |- | |''Brachycope'' {{small|Reichenow, 1900}} | * [[Bob-tailed weaver]] (''Brachycope anomala'') |- |[[File:Euplectes progne male South Africa cropped.jpg|175px]] |''[[Euplectes]]'' {{small|Swainson, 1829}} | * [[Yellow-crowned bishop]] (''Euplectes afer'') * [[Fire-fronted bishop]] (''Euplectes diadematus'') * [[Black bishop]] (''Euplectes gierowii'') * [[Black-winged red bishop]] (''Euplectes hordeaceus'') * [[Northern red bishop]] or orange bishop (''Euplectes franciscanus'') * [[Southern red bishop]] or red bishop (''Euplectes orix'') * [[Zanzibar red bishop]] (''Euplectes nigroventris'') * [[Golden-backed bishop]] (''Euplectes aureus'') * [[Yellow bishop]] (''Euplectes capensis'') * [[Fan-tailed widowbird]] (''Euplectes axillaris'') * [[Yellow-mantled widowbird]] (''Euplectes macroura'') * [[White-winged widowbird]] (''Euplectes albonotatus'') * [[Red-collared widowbird]] (''Euplectes ardens'') * [[Red-cowled widowbird]] (''Euplectes laticauda'') * [[Marsh widowbird]] (''Euplectes hartlaubi'') * [[Montane widowbird]] (''Euplectes psammacromius'') * [[Long-tailed widowbird]] (''Euplectes progne'') * [[Jackson's widowbird]] (''Euplectes jacksoni'') |- |}
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