Editing Trachypithecus (section)

== Evolution ==
[[File:Trachypithecus cristatus robustus.jpg|thumb|''Trachypithecus cristatus robustus'' skull]]

Genetic analysis indicates that the ancestors of the modern species of lutung first differentiated from one another a little over 3 million years ago, during the late [[Pliocene]]. The various species alive today then diverged during the [[Pleistocene]], presumably driven by habitat changes during the [[Ice age|Ice Ages]].<ref name=Roos2008>{{cite journal | author = Roos, C.| year = 2008 | title = Mitochondrial phylogeny, taxonomy and biogeography of the silvered langur species group (''Trachypithecus cristatus'') | journal = Molecular Phylogenetics and Evolution | volume = 47 | issue = 2 | pages = 629–636 | doi = 10.1016/j.ympev.2008.03.006 | pmid=18406631| bibcode = 2008MolPE..47..629R |display-authors=etal}}{{dead link|date=March 2019|bot=medic}}{{cbignore|bot=medic}}</ref> The oldest fossils clearly identified as belonging to the genus date from the middle [[Pleistocene]] of [[Vietnam]] and [[Laos]]; later fossils are also known from Thailand, Java, and Sumatra. The closest living relatives of the lutungs are probably either the [[gray langur]]s or the surilis, although the exact relationships remain unclear, possibly due to hybridisation between these genera during the course of their recent evolutionary history.<ref name=Harding2010>{{cite journal | author = Harding, L.E. | year = 2010 | title = ''Trachypithecus cristatus'' (Primates: Cercopithecidae) | journal = Mammalian Species | volume = 42 | issue = 1 | pages = 149–165 | doi = 10.1644/862.1 | doi-access = free }}</ref>