Editing Northern crested newt (section)

==Evolution==

{{Annotated image
| image = Wielstra2019 Triturus Graphical Abstract blank.jpg
| width = 300
| image-width = 300
| float = right
| annotations =
{{Annotation|34|28|marbled<br />newts|font-size=13}}
{{Annotation|2|155|crested<br />newts|font-size=13|text-align=right}}
{{Annotation|140|40|''[[Triturus marmoratus|T. marmoratus]]'' and<br />''[[Triturus pygmaeus|T. pygmaeus]]''|font-size=13}}
{{Annotation|140|110|''[[Triturus karelinii|T. karelinii]]'', ''[[Triturus anatolicus|T. anatolicus]]''<br />and ''[[Triturus ivanbureschi|T. ivanbureschi]]''|font-size=13}}
{{Annotation|140|180|''[[Triturus carnifex|T. carnifex]]'' and<br />''[[Triturus macedonicus|T. macedonicus]]''|font-size=13}}
{{Annotation|140|250|'''''T. cristatus'''''|font-size=13}}
{{Annotation|140|310|''[[Triturus dobrogicus|T. dobrogicus]]''|font-size=13}}
| caption = Position of the northern crested newt (''T. cristatus'') in the [[phylogenetic tree]] of the genus ''Triturus''{{r|WielstraMcCartney-Melstad2019}}
}}

The northern crested newt sometimes hybridises with other crested newt species where their ranges meet, but overall, the different species are [[reproductive isolation|reproductively isolated]].{{r|ArntzenWielstra2014}} In a case study in the Netherlands, genes of the introduced Italian crested newt (''T. carnifex'') were found to [[introgression|introgress]] into the [[gene pool]] of the native northern crested newt.{{r|MeilinkArntzen2015}} The closest relative of the northern crested newt, according to [[molecular phylogenetic]] analyses, is the Danube crested newt (''T. dobrogicus'').{{r|WielstraMcCartney-Melstad2019|WielstraArntzen2011}}

In western France, the northern crested newt's range overlaps with that of the marbled newt (''T. marmoratus''), but the two species in general prefer different habitats.{{r|Zuiderwijk2004|SchoorlZuiderwijk1980}} When they do occur in the same breeding ponds, they can form hybrids, which have intermediate characteristics. Hybrids resulting from the cross of a crested newt male with a marbled newt female are much rarer due to increased mortality of the larvae and consist only of males. In the reverse cross, males have lower survival rates than females. Overall, viability is reduced in these hybrids and they rarely [[Backcrossing|backcross]] with their parent species. Hybrids made up 3–7% of the adult populations in different studies.{{r|ArntzenJehle2009}}

Little [[genetic variation]] was found over most of the species' range, except in the [[Carpathians]]. This suggests that the Carpathians was a [[refugium (population biology)|refugium]] during the [[Last Glacial Maximum]]. The northern crested newt then expanded its range north-, east- and westwards when the climate rewarmed.{{r|BabikPabijan2009|WielstraBabik2015|WielstraZielinski2017}}