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Place cell
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====Olfactory inputs==== Although place cells primarily rely on visuospatial input, some studies suggest that olfactory input may also affect the formation and stability of place fields.<ref name=Save2000>{{cite journal|last=Save|first=Etienne|author2=Ludek Nerad |author3=Bruno Poucet |title=Contribution of multiple sensory information to place field stability in hippocampal place cells|journal=Hippocampus|date=23 February 2000|volume=10|issue=1|pages=64β76|doi=10.1002/(SICI)1098-1063(2000)10:1<64::AID-HIPO7>3.0.CO;2-Y|pmid=10706218|s2cid=34908637 }}</ref><ref>{{Cite journal|last1=Poucet|first1=Bruno|last2=Save|first2=Etienne|last3=Lenck-Santini|first3=Pierre-Pascal|date=2011|title=Sensory and Memory Properties of Hippocampal Place Cells|journal=Reviews in the Neurosciences|volume=11|issue=2β3|pages=95β112|doi=10.1515/REVNEURO.2000.11.2-3.95|pmid=10718148|s2cid=1952601|issn=2191-0200}}</ref><ref name="Jeffery-2003">{{Cite book|url=https://books.google.com/books?id=ZMLADwAAQBAJ&q=olfaction+%22place+cells%22&pg=PA48|title=The Neurobiology of Spatial Behaviour|last=Jeffery|first=Kathryn J.|date=2003|publisher=Oxford University Press|isbn=978-0-19-851524-1|language=en}}</ref> Olfaction may compensate for a loss of visual information,<ref name="Save2000" /><ref name="Jeffery-2003" /> or even be responsible for the formation of stable place fields in the same way visuospatial cues are.<ref name="Zhang 2013">{{cite journal|last=Zhang|first=Sijie|author2=Denise Manahan-Vaughn|date=5 September 2013|title=Spatial Olfactory Learning Contributes to Place Field Formation in the Hippocampus|journal=Cerebral Cortex|volume=25|issue=2|pages=423β432|doi=10.1093/cercor/bht239|pmc=4380081|pmid=24008582}}</ref> This has been confirmed by a study in a virtual environment that was composed of odor gradients.<ref name="Radvansky 2018">{{cite journal|last=Radvansky|first=Brad|author2=Daniel Dombeck|date=26 February 2018|title=An olfactory virtual reality system for mice|journal=Nature Communications|volume=9|issue=1|page=839|doi=10.1038/s41467-018-03262-4|pmc=5827522|pmid=29483530|bibcode=2018NatCo...9..839R}}</ref> Change in the olfactory stimulus in an environment may also cause the remapping of place cells.<ref name="Anderson-2003" /><ref name="Jeffery 775β785" />
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