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===Osmoregulation === In sea turtles, the bladder is one unit and in most freshwater turtles, it is double-lobed.<ref>{{cite book |last1=Miller |first1=Jeffrey D. |last2=Dinkelacker|first2=Stephen A.|chapter=Reproductive Structures and Strategies of Turtles |editor-last=Wyneken |editor-first=Jeanette |editor-last2=Bels |editor-first2=V. L. |editor-last3=Godfrey |editor-first3=Matthew H. |title=Biology of Turtles |year=2008 |publisher=CRC Press |isbn=978-0-8493-3339-2 |oclc=144570900 |page=234}}</ref> Sea turtle bladders are connected to two small accessory bladders, located at the sides to the neck of the urinary bladder and above the [[Pubis (bone)|pubis]].<ref>{{cite journal |last1=Wyneken |first1=Jeanette |last2=Witherington |first2=Dawn |date=February 2015 |title=Urogenital System |url=http://www.ivis.org/advances/wyneken/16.pdf?LA |journal=Anatomy of Sea Turtles |volume=1 |pages=153β165 |access-date=May 18, 2021 |archive-date=June 8, 2019 |archive-url=https://web.archive.org/web/20190608150657/http://www.ivis.org/advances/wyneken/16.pdf?LA |url-status=live }}</ref> Arid-living tortoises have bladders that serve as reserves of water, storing up to 20% of their body weight in fluids. The fluids are normally low in [[solute]]s, but higher during droughts when the reptile gains [[potassium]] salts from its plant diet. The bladder stores these salts until the tortoise finds fresh drinking water.<ref name="Bentley 2013">{{cite book |last=Bentley |first=Peter J. |url=https://books.google.com/books?id=U0D3BwAAQBAJ&pg=PA143 |title=Endocrines and Osmoregulation: A Comparative Account in Vertebrates |date=2013 |publisher=Springer |page=143 |isbn=978-3-662-05014-9 |access-date=May 18, 2021 |archive-date=May 31, 2021 |archive-url=https://web.archive.org/web/20210531050224/https://books.google.com/books?id=U0D3BwAAQBAJ&pg=PA143 |url-status=live }}</ref> To regulate the amount of salt in their bodies, sea turtles and the [[brackish water|brackish]]-living [[diamondback terrapin]] secrete excess salt in a thick sticky substance from their [[lacrimal gland|tear gland]]s. Because of this, sea turtles may appear to be "crying" when on land.{{sfn|Franklin|2011|p=31}}
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