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Tooth enamel
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== Other organisms == Enamel or [[enameloid]] is found in the [[dermal denticle]]s of [[shark]]s and many early vertebrates,<ref>{{cite journal |last1=Mondéjar-Fernández |first1=Jorge |last2=Meunier |first2=François J. |last3=Cloutier |first3=Richard |last4=Clément |first4=Gaël |last5=Laurin |first5=Michel |title=A microanatomical and histological study of the scales of the Devonian sarcopterygian Miguashaia bureaui and the evolution of the squamation in coelacanths |journal=Journal of Anatomy |date=2021 |volume=239 |issue=2 |pages=451–478 |doi=10.1111/joa.13428 |pmid=33748974 |pmc=8273612 |url=https://doi.org/10.1111/joa.13428 |language=en |issn=1469-7580}}</ref> and it appeared there before [[Gnathostomata|gnathostome]] [[Tooth|teeth]] evolved.<ref>{{cite journal |last1=Mondéjar-Fernández |first1=Jorge |last2=Janvier |first2=Philippe |title=Finned Vertebrates |journal=Vertebrate Skeletal Histology and Paleohistology |date=2021 |pages=294–324 |doi=10.1201/9781351189590-15 |url=https://www.taylorfrancis.com/chapters/edit/10.1201/9781351189590-15/finned-vertebrates-jorge-mond%C3%A9jar-fern%C3%A1ndez-philippe-janvier |publisher=CRC Press|isbn=978-1-351-18959-0 |s2cid=241700775 |url-access=subscription }}</ref> The [[Ganoine|ganoin]] that covers the scales of many [[Actinopterygii|actinopterygians]] is probably derived from enamel.<ref>{{cite journal |last1=Zylberberg |first1=Louise |last2=Meunier |first2=François |last3=Laurin |first3=Michel |title=A microanatomical and histological study of the postcranial dermal skeleton of the Devonian actinopterygian Cheirolepis canadensis |journal=Acta Palaeontologica Polonica |date=2015 |doi=10.4202/app.00161.2015 |s2cid=55113048 |language=en |issn=0567-7920|doi-access=free }}</ref> Enamel-like substances also coat the jaws of some crustacea, but this is not homologous with vertebrate enamel.<ref name="ref_">{{Cite journal | doi = 10.1038/485419a | title = Evolution: Tooth-enamel similarities | journal = Nature | volume = 485 | issue = 7399 | pages = 419 | year = 2012 | bibcode = 2012Natur.485Q.419. | s2cid = 52798363 | doi-access = free }}</ref><ref name="Bentov2012">{{Cite journal | last1 = Bentov | first1 = S. | last2 = Zaslansky | first2 = P. | last3 = Al-Sawalmih | first3 = A. | last4 = Masic | first4 = A. | last5 = Fratzl | first5 = P. | last6 = Sagi | first6 = A. | last7 = Berman | first7 = A. | last8 = Aichmayer | first8 = B. | doi = 10.1038/ncomms1839 | title = Enamel-like apatite crown covering amorphous mineral in a crayfish mandible | journal = Nature Communications | volume = 3 | pages = 839 | year = 2012 | pmid = 22588301| pmc = 3382302| issue=5| bibcode = 2012NatCo...3..839B }}</ref> [[Enameloid]] covers some fish scales.
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