Editing Toe (section)

==Other animals==
[[File:Pakicetidae_-_Pakicetus_attocki.JPG|thumb|The skeleton of ''[[Pakicetus]]'', an extinct digitigrade mammal.]]
In [[bird]]s with [[anisodactyl]] or [[heterodactyl]] feet, the hallux is opposed or directed backwards and allows for grasping and perching.

While the thumb is often mentioned{{by whom?|date=August 2019}} as one of the signature characteristics in humans, this manual digit remains partially primitive and is actually present in all [[primates]]. In humans, the most [[Synapomorphy |derived]] digital feature is the hallux.<ref>{{cite journal
| last1 = Lovejoy | first1 = C. Owen
| last2 = Suwa | first2 = Gen
| last3 = Simpson | first3 = Scott W.
| last4 = Matternes | first4 = Jay H.
| last5 = White | first5 = Tim D.
| title = The Great Divides: Ardipithecus ramidus Reveals the Postcrania of Our Last Common Ancestors with African Apes
| journal = Science |date=October 2009 | volume = 326 | issue = 5949 | pages = 100–6
| doi = 10.1126/science.1175833
| url = https://www.science.org/doi/abs/10.1126/science.1175833
| pmid = 19810199
| s2cid = 19629241 | url-access = subscription}}
</ref>