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Asplenium
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==Taxonomy and genetics== Many groups of species have been separated from ''Asplenium'' as [[Segregate (taxonomy)|segregate]] genera. These include ''Camptosorus'', ''Ceterach'', ''Phyllitis'', and ''Tarachia'', but these species can form [[hybrid (biology)|hybrid]]s with other ''Asplenium'' species and because of this are usually included in a more broadly defined ''Asplenium''.<ref>{{cite journal | last1=Schneider | first1=Harald | last2=Russell | first2=Steve J. | last3=Cox | first3=Cymon J. | last4=Bakker | first4=Freek | last5=Henderson | first5=Sally | last6=Rumsey | first6=Fred | last7=Barrett | first7=John | last8=Gibby | first8=Mary|author-link8=Mary Gibby| last9=Vogel | first9=Johannes C. | author-link9=Johannes Vogel (botanist) | title=Chloroplast Phylogeny of Asplenioid Ferns based on rbcL and trnL-F Spacer Sequences (Polypodiidae, Aspleniaceae) and its Implications for Biogeography | journal=[[Systematic Botany]] | volume=29 | issue=2 | year=2004 | pages=260β274 | doi=10.1600/036364404774195476 | bibcode=2004SysBo..29..260S | s2cid=85868809 | ref = {{harvid|Schneider et al.|2004}}}}</ref> Some of the older classifications elevate the Aspleniaceae to the [[taxonomic rank]] of [[order (biology)|order]] as Aspleniales. The newer classifications place it in the subordinal group called eupolypods within the order [[Polypodiales]]. Within the eupolypods, Aspleniaceae belongs to a [[clade]] informally and provisionally known as eupolypods II. It has been found that in some species, the [[chloroplast]] [[plastome|genome]] has [[Introduction to evolution|evolved]] in complex and highly unusual ways. This makes standard [[cladistic]] analyses unsuited to resolve the [[phylogeny]] of that particular group of ferns, and even very sophisticated [[computational phylogenetics]] methods yield little information. In addition to hybridization running rampant in parts of this genus, there are also some species like the [[mother spleenwort]] (''A. bulbiferum'') or ''[[Asplenium viviparum|A. viviparum]]'' which mainly [[asexual reproduction|reproduce asexually]], essentially [[cloning]] themselves over and over again. While most are [[diploid]] or [[tetraploid]], some species (e.g. ''[[Asplenium shuttleworthianum|A. shuttleworthianum]]'') are octoploid.<ref>{{cite journal |author=Lara D. Shepherd, Barbara R. Holland & Leon R. Perrie |year=2008 |title=Conflict amongst chloroplast DNA sequences obscures the phylogeny of a group of ''Asplenium'' ferns |journal=[[Molecular Phylogenetics and Evolution]] |volume=48 |issue=1 |pages=176β187 |doi=10.1016/j.ympev.2008.02.023 |pmid=18462954|bibcode=2008MolPE..48..176S }}</ref>
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