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== Taxonomy and systematics == [[File:Regulus ignicapillus Roitelet triple-bandeau.jpg|thumb|A temporarily stunned adult male found on a pavement in [[Lille]], France. The pattern on its head is seen clearly.]] The kinglets are a small group of birds sometimes included in the [[Old World warbler]]s, but frequently given family status,<ref name = BB852 >{{cite journal |last= Monroe |first= Burt L. |date=February 1992 |title= The new DNA-DNA avian classification: What's it all about? |journal= [[British Birds (magazine)|British Birds]] |volume=85 |issue= 2|pages=53–61 }}</ref> especially as recent research shows that, despite superficial similarities, the crests are [[phylogenetic]]ally remote from the warblers.<ref name= Barker>{{cite journal | last= Barker | first= F Keith | author2=Barrowclough, George F|author3=Groth, Jeff G | year = 2002 | title= A phylogenetic hypothesis for passerine birds: taxonomic and biogeographic implications of an analysis of nuclear DNA sequence data | journal= [[Proceedings of the Royal Society of London B]] | volume= 269 | pages= 295–308 | doi = 10.1098/rspb.2001.1883 | pmid=11839199 | pmc=1690884 | issue=1488}}</ref><ref name= Spicer>{{cite journal | last= Spicer | first= Greg S |author2=Dunipace, Leslie | year= 2004 | title= Molecular phylogeny of songbirds (Passeriformes) inferred from mitochondrial 16S ribosomal RNA gene sequences | journal= Molecular Phylogenetics and Evolution | volume= 30 | issue= 2 | pages= 325–335 | url = http://online.sfsu.edu/~gs/spicer/pages/spicerpdf/spicerdunipace04.pdf| doi = 10.1016/S1055-7903(03)00193-3 | pmid=14715224}}</ref><ref name= alstrom>{{cite journal | last= Alström | first= Per | author2= Ericson, Per G P | author3= Olsson, Urban | author4= Sundberg, Per | year= 2006 | title= Phylogeny and classification of the avian superfamily Sylvioidea | journal= Molecular Phylogenetics and Evolution | volume= 38 | issue= 2 | pages= 381–397 | url= http://www.nrm.se/download/18.4e1d3ca810c24ddc70380001143/Alstr%C3%B6m+et+al+Sylvioidea+MPEV+2006.pdf | doi= 10.1016/j.ympev.2005.05.015 | pmid= 16054402 | access-date= 2010-11-24 | archive-date= 2021-06-27 | archive-url= https://web.archive.org/web/20210627134311/https://www.nrm.se/download/18.4e1d3ca810c24ddc70380001143/Alstr%C3%B6m+et+al+Sylvioidea+MPEV+2006.pdf | url-status= dead }}</ref> The names of the family [[Regulidae]], and the genus ''[[kinglet|Regulus]]'', are derived from the [[Latin]] ''regulus'', a diminutive of ''rex'', "a king",<ref name = Chambers>{{cite book | title = The Chambers Dictionary | edition = 9th | year =2006 | publisher = Chambers | location = Edinburgh | pages = 223, 735, 1277 | isbn = 978-0-550-10185-3| title-link = Chambers Dictionary }}</ref> and refer to the characteristic orange or yellow crests of adult kinglets. The common firecrest was first formally [[species description|described]] by Dutch zoologist [[Coenraad Jacob Temminck]] in 1820 as ''Sylvia ignicapilla'';<ref name= Temminck >{{cite book | last = Temminck | first = Coenraad Jacob | author-link = Coenraad Jacob Temminck| title = Manuel d'ornithologie, ou Tableau systematique des oiseaux qui se trouvent en Europe | edition= second | year = 1820–1840 | publisher = H Cousin & E d'Ocagne | location = Paris| page = 231 |url=https://books.google.com/books?id=BN4TAAAAYAAJ&q=Manuel%20d'ornithologie%2C%20ou%20Tableau%20systematique%20des%20oiseaux%20qui%20se%20trouvent%20en%20Europe&pg=PA231 | language = fr}}</ref> the relatively late identification of this common European bird arose from a perception that it was just a variety of the goldcrest.<ref name= Wood>{{cite book | last = Wood | first = Neville | year = 1836 | title = British song birds: being popular descriptions and anecdotes of the choristers of the groves | location = London| publisher = John W Parker | pages = [https://archive.org/details/britishsongbirds00wood/page/143 143]–145 | url = https://archive.org/details/britishsongbirds00wood| quote = British song birds: being popular descriptions and anecdotes of the choristers of the groves. }}</ref> The species name is derived from Latin ''ignis'' "fire" and ''capillus'' "hair".<ref name = Chambers/> The binomial is frequently given as ''R. ignicapillus'' due to a misunderstanding of Latin grammar.<ref name= aerc2010>{{cite book | author = Crochet, P-A | author2 = Raty, L | author3 = De Smet, G | author4 = Anderson, B | author5 = Barthel, P H | author6 = Collinson, J M | author7 = Dubois, P J | author8 = Helbig, A J | author9 = Jiguet, F | author10 = Jirle, E | author11 = Knox, A G | author12 = Le Maréchal, P | author13 = Parkin, D T | author14 = Pons, J-M | author15 = Roselaar, C S | author16 = Svensson, L | author17 = van Loon, A J | author18 = Yésou, P | title = AERC TAC's Taxonomic Recommendations July 2010 | url = http://www.aerc.eu/DOCS/AERC%20TAC%20recommendations%20July%202010%20version%202.0.pdf | year = 2010 | publisher = Association of European Rarity Committees (AERC) | page = 14 }}</ref> There are two widely recognised subspecies of common firecrest, nominate ''R. i. ignicapilla'' and Mediterranean ''R. i. balearicus'' (Jordans, 1923). The latter form is found on the Balearic Islands and in north Africa, and is slightly paler below and greyer above than the nominate subspecies.<ref name= Baker/> Other subspecies have been claimed, including southeastern ''R. i. caucasicus'', North African ''R. i. laeneni'',<ref name="Packert"/> and [[Crimea]]n ''R. i. tauricus''.<ref name= Redkin>{{cite journal | last= Redkin | first= Y A| year= 2001| title= A new subspecies of Firecrest ''Regulus ignicapillus'' (Temminck, 1820) (Regulidae, Passeriformes) from the mountains of the Crimea | journal= Ornitologia | volume= 29 | pages= 98–102 }}</ref> The [[Madeira firecrest]], ''R. madeirensis'', was formerly also considered to be a subspecies of the common firecrest, but [[Phylogenetics|phylogenetic]] analysis based on the [[cytochrome b|cytochrome ''b'' gene]] showed that the Madeiran form is distinct at the species level. Cytochrome ''b'' gene divergence between the Madeira firecrest and the European bird is 8.5%, comparable with the divergence level between other recognised ''Regulus'' species, such as the 9% between the goldcrest and the [[golden-crowned kinglet]].<ref name = hbw>Martens, Jochen; Päckert, Martin "Family Regulidae (Kinglets & Firecrests)" pp. 330–349 in {{cite book| editor1-last = Del Hoyo | editor1-first = Josep | editor2-last = Elliott|editor2-first=Andrew|editor3-last=Christie|editor3-first=David A | title = Handbook of the Birds of the World: Old World Flycatchers to Old World Warblers v. 11 | year = 2006 | publisher = [[Lynx Edicions]] | location = Barcelona | isbn = 978-84-96553-06-4 | title-link = Handbook of the Birds of the World }}</ref> The island form also differs in [[morphology (biology)|morphology]] and vocalisations.<ref name= Sangster>{{cite journal | last= Sangster | first= George | author-link = George Sangster | author2= Collinson, J Martin| author3=Helbig, Andreas J| author4=Knox, Alan G| author5=Parkin, David T. | year= 2005| title= Taxonomic recommendations for British birds: third report | journal= [[Ibis (journal)|Ibis]] | volume= 147 | issue= 4 | pages = 821–826 | doi =10.1111/j.1474-919X.2005.00483.x| s2cid= 250043960 | doi-access= free }}</ref> The proposed split was accepted by the [[Association of European Rarities Committees]] (AERC) in 2003,<ref name= aerc >{{cite book | author = AERC Taxonomy Committee | title = AERC TAC's Taxonomic Recommendations | url = http://www.aerc.eu/DOCS/AERCTACAccepted.pdf | year = 2003| publisher = [[Association of European Rarities Committees]] | page = 22}}</ref> with most other authorities also splitting it later on. The [[flamecrest]] or Taiwan firecrest (''Regulus goodfellowi'') of Taiwan has sometimes been viewed as a race of the common firecrest; however, the flamecrest's territorial song, which resembles those of the [[Himalayas|Himalayan]] races of the goldcrest, and genetic data indicate that the flamecrest is closely related to the Himalayan goldcrest and only distantly to the two firecrest species.<ref name= flamecrest>{{cite journal | last= Päckert | first= Martin |author2=Martens, Jochen|author3=Severinghaus, Lucia Liu | year=2008 | title= The Taiwan Firecrest (''Regulus goodfellowi'') belongs to the Goldcrest assemblage (''Regulus regulus'' s. l.): evidence from mitochondrial DNA and the territorial song of the Regulidae | journal= Journal of Ornithology | volume= 150 | issue = 1 | pages= 205–220 | doi = 10.1007/s10336-008-0335-5 | s2cid= 5626256 }}</ref> The kinglets on the [[Canary Islands]], which were also considered to be close to firecrests, have now been shown to comprise two subspecies of goldcrest.<ref name= pack2006>{{cite journal | last= Päckert | first= Martin | year=2006 | title= Song dialects as diagnostic characters—acoustic differentiation of the Canary Island Goldcrest subspecies ''Regulus regulus teneriffae'' Seebohm 1883 and ''R. r. ellenthalerae'' Päckert et al. 2006 (Aves: Passeriformes: Regulidae) | journal= Zootaxa | volume= 1325 | pages= 99–115| doi= 10.11646/zootaxa.1325.1.7 }}</ref> === Fossils === There are a few [[Pleistocene]] (2.6 million to 12,000 years ago) records from Europe and Israel of extant ''Regulus'' species, mostly goldcrests or unidentifiable to species, but also a Spanish specimen of firecrest. A left [[ulna]] from Bulgaria was identified as belonging to a fossil species, ''[[Regulus bulgaricus]]'', from 2.6–1.95 [[mya (unit)|mya]]. This appears to be ancestral to the common firecrest, with the goldcrest diverging from this lineage in the Middle Pleistocene.<ref name=Boev>{{cite journal|last=Boev |first=Zlatozar |year=1999 |title=''Regulus bulgaricus'' sp. n. – the first fossil Kinglet (Aves: Sylviidae) from the Late Pliocene of Varshets, Western Bulgaria |journal=Historia Naturalis Bulgarica |volume=10 |url=https://www.biodiversitylibrary.org/page/31093140#page/111/mode/1up |pages=109–115}}</ref>
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