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Formiciinae
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==Size== While workers belonging to the subfamily have not been found, queens and males for ''T. giganteum'' and ''T. simillimum'' are known and ''T. lubei'' is known from a single queen. The average size for the queens and males in ''Titanomyrma'' is equal to that of some of the largest modern ants known. Only the queens of ''[[Dorylus wilverthi]]'' currently reach similar lengths, up to {{convert|52|mm|in}} as the smallest species of ''Titanomyrma'', ''T. lubei''.<ref name="Archibald2011"/> Formiciinae members were restricted in habitat to living in regions which had a [[mesic habitat|mesic]] wet climate and an average mean annual temperature of {{convert|20|Β°C|Β°F}} or higher. This is similar to the restricted ranges of the largest species of modern ants.<ref name="Archibald2011"/> The spread of the subfamily from Europe to North America is postulated to have been across the [[North Atlantic]] landbridges which were present in the Eocene. While the average temperatures for this route are thought to have been lower than the range needed for Formiciinae species, a series of warmer events throughout the Eocene are suggested as aides in the crossing.<ref name="Archibald2011"/>
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