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Huperzia
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==Taxonomy== The genus ''Huperzia'' was created by [[Johann Jakob Bernhardi]] in 1801. Bernhardi separated ''Huperzia'' from ''Lycopodium''. The [[type species]] is ''Lycopodium selago'' which became ''[[Huperzia selago]]''.<ref name=IPNI_30000333-2/> In the [[Pteridophyte Phylogeny Group]] classification of 2016 (PPG I), ''Huperzia'' is placed in the subfamily [[Huperzioideae]] of the family [[Lycopodiaceae]].<ref name=PPGI/> A [[Phylogenetics|phylogenetic]] study in 2016, employing both molecular and morphological data, concluded that either a one-genus or a three-genus division of the subfamily produced [[Monophyly|monophyletic]] taxa. The authors preferred the three-genus division, recognizing ''Huperzia'', ''[[Phlegmariurus]]'' and ''[[Phylloglossum]]''. Their preferred hypothesis for the relationships of the three genera was:<ref name=FielTestBostHolt16/> {{clade |label1=[[Huperzioideae]] |1={{clade |1={{clade |1=''[[Phylloglossum]]'' |2=''Huperzia'' }} |2=''[[Phlegmariurus]]'' }} }} The majority of the species formerly placed in a broadly defined ''Huperzia'' belong in ''Phlegmariurus''.<ref name=FielTestBostHolt16/> Earlier, the ''Flora of North America'' had also separated ''Huperzia'' from ''Phegmariurus''.<ref name=FNA/> However, ''Phlegmariurus'' is difficult to separate morphologically, and others have preferred the one-genus division of the subfamily.<ref name=CFLW_Phl/><ref name=POWO_30000333-2/> ===Species=== The PPG I classification stated there were 25 species in the genus ''Huperzia''.<ref name=PPGI/> {{As of|2024|June}}, ''World Ferns'' listed the following species, noting that "many species still need transfer into other split genera".<ref name=CFLW_Hup/> {{Div col|colwidth=25em}} *''[[Huperzia acicularis]]'' <small>Björk</small>, provisionally accepted *''[[Huperzia appalachiana]]'' <small>Beitel & Mickel</small> *''[[Huperzia archboldiana]]'' <small>(Nessel) Holub</small> *''[[Huperzia arunachalensis]]'' <small>(D.D.Pant & P.S.Pandey) Fraser-Jenk.</small> *''[[Huperzia asiatica]]'' <small>(Ching) N.Shrestha & X.C.Zhang</small> *''[[Huperzia australiana]]'' <small>(Herter) Holub</small> *''[[Huperzia beccarii]]'' <small>(Alderw.) Holub</small> *''[[Huperzia beiteliana]]'' <small>Mickel</small> *''[[Huperzia bucahwangensis]]'' <small>Ching</small> *''[[Huperzia campestris]]'' <small>(Alderw.) Holub</small> *''[[Huperzia catharinae]]'' <small>(Christ) Holub</small> *''[[Huperzia ceylanica]]'' <small>(Spring) Rothm.</small> *''[[Huperzia chinensis]]'' <small>(Christ) Ching</small> *''[[Huperzia chishuiensis]]'' <small>X.Y.Wang & P.S.Wang</small> *''[[Huperzia continentalis]]'' <small>Testo, A.Haines & A.V.Gilman</small> *''[[Huperzia crispata]]'' <small>(Ching) Ching</small> *''[[Huperzia delavayi]]'' <small>(Christ & Herter) Ching</small> *''[[Huperzia emeiensis]]'' <small>(Ching & H.S.Kung) Ching & H.S.Kung</small> *''[[Huperzia erosa]]'' <small>Beitel & W.H.Wagner</small> *''[[Huperzia erubescens]]'' <small>(Brack.) Holub</small> *''[[Huperzia europaea]]'' <small>Björk</small>, provisionally accepted *''[[Huperzia everettii]]'' <small>(Herter) Holub</small> *''[[Huperzia fuegiana]]'' <small>(Roiv.) Holub</small> *''[[Huperzia gedeana]]'' <small>(Alderw.) Holub</small> *''[[Huperzia goliathensis]]'' <small>(Alderw.) Holub</small> *''[[Huperzia haleakalae]]'' <small>(Brack.) Holub</small> *''[[Huperzia herteriana]]'' <small>(Kümmerle) T.Sen & U.Sen</small> *''[[Huperzia javanica]]'' <small>(Sw.) Fraser-Jenk.</small> *''[[Huperzia jejuensis]]'' <small>B.Y.Sun & J.Lim</small> *''[[Huperzia kangdingensis]]'' <small>(Ching) Ching</small> *''[[Huperzia kunmingensis]]'' <small>Ching</small> *''[[Huperzia laipoensis]]'' <small>Ching</small> *''[[Huperzia lajouensis]]'' <small>Ching</small> *''[[Huperzia leishanensis]]'' <small>X.Y.Wang</small> *''[[Huperzia liangshanica]]'' <small>(H.S.Kung) Ching & H.S.Kung</small> *''[[Huperzia lucidula]]'' <small>(Michx.) Trevis.</small> *''[[Huperzia medogensis]]'' <small>Ching & Y.X.Lin</small> *''[[Huperzia meghalaica]]'' <small>Fraser-Jenk.</small> *''[[Huperzia miniata]]'' <small>(Spring) Trevis.</small> *''[[Huperzia minima]]'' <small>(Herter) Holub</small> *''[[Huperzia miyoshiana]]'' <small>(Makino) Ching</small> *''[[Huperzia muscicola]]'' <small>W.M.Chu</small> *''[[Huperzia myriophyllifolia]]'' <small>(Hayata) Holub</small> *''[[Huperzia nanchuanensis]]'' <small>(Ching & H.S.Kung) Ching & H.S.Kung</small> *''[[Huperzia nanlingensis]]'' <small>Shrestha, F.W.Xing, X.P.Qi, Y.H.Yan & X.C.Zhang</small> *''[[Huperzia occidentalis]]'' <small>(Clute) Kartesz & Gandhi</small> *''[[Huperzia porophila]]'' <small>(F.E.Lloyd & Underw.) Holub</small> *''[[Huperzia quasipolytrichoides]]'' <small>(Hayata) Ching</small> *''[[Huperzia rubicaulis]]'' <small>S.K.Wu & X.Cheng</small> *''[[Huperzia saururoides]]'' <small>(Bory & D'Urv.) Rothm.</small> *''[[Huperzia selago]]'' <small>(L.) Bernh.</small> (including [[Huperzia appressa|''H. selago'' subsp. ''appressa'']] - Appalachian firmoss) *''[[Huperzia serrata]]'' <small>(Thunb.) Trevis.</small> *''[[Huperzia shresthae]]'' <small>Fraser-Jenk.</small> *''[[Huperzia somae]]'' <small>(Hayata) Ching</small> *''[[Huperzia sprengeri]]'' <small>(Nessel) Holub</small> *''[[Huperzia suberecta]]'' <small>(Lowe) Tardieu</small> *''[[Huperzia subintegra]]'' <small>(Hillebr.) Beitel & W.H.Wagner</small> *''[[Huperzia sumatrana]]'' <small>(Alderw.) Holub</small> *''[[Huperzia sutchueniana]]'' <small>(Herter) Ching</small> *''[[Huperzia tibetica]]'' <small>(Ching) Ching</small> *''[[Huperzia yakusimensis]]'' <small>(Herter) Holub</small> *''[[Huperzia zollingeri]]'' <small>(Herter) Holub</small> {{Div col end}} The following hybrids have been described:<ref name=CFLW_Hup/> *[[Huperzia × bartleyi|''Huperzia'' × ''bartleyi'']] <small>(Cusick) Holub</small> *[[Huperzia × buttersii|''Huperzia'' × ''buttersii'']] <small>(Abbe) Kartesz & Gandhi</small> *[[Huperzia × carlquistii|''Huperzia'' × ''carlquistii'']] <small>Beitel & W.H.Wagner</small> *[[Huperzia × gillettii|''Huperzia'' × ''gillettii'']] <small>Beitel & W.H.Wagner</small> *[[Huperzia × josephbeitelii|''Huperzia'' × ''josephbeitelii'']] <small>A.Haines</small> *[[Huperzia × medeirosii|''Huperzia'' × ''medeirosii'']] <small>Beitel & W.H.Wagner</small> *[[Huperzia × protoporophila|''Huperzia'' × ''protoporophila'']] <small>A.Haines</small> *[[Huperzia × sulcinervia|''Huperzia'' × ''sulcinervia'']] <small>(Spring) Trevis.</small>
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