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Lagoon triggerfish
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==Mating and reproduction== Both sexes guard territories, some maintaining a territory for eight years or longer (with males holding territories for significantly longer than females). A typical male territory may overlap with one to five female territories, and their mating system is described as haremic, although not much is known about this (similar mating systems are seen in other [[Balistidae]] species). If a male or female is removed or disappears their territories are soon taken over by a new fish. They reproduce multiple times over their lifetimes.<ref name="Kuwamura" /> Pair-spawning takes place around sunrise, with the egg masses being attached to sand, coral rubble or algae. They hatch on the same day around sunset. Although paternal care is normal in teleost fishes with external fertilization, it is the mothers in this species that guard and care for eggs until they hatch. The mother remains above the eggs for about 12β14 hours, fanning the eggs with her pectoral fins to improve aeration for perhaps 30% of the time. She chases away most fish that approach and remove other intruders like starfish by mouth. Maternal care is effective in preventing predation, and experimental removal of the mothers reduced survival to almost nothing suggesting this behaviour is adaptive. Unlike fathers, mothers forage less and over a smaller area near the egg mass while caring for the eggs. Since the males have multiple mates, caring for an egg mass would probably be more costly in terms of lost mating opportunities so maternal care is considered to be an [[evolutionarily stable strategy]].<ref name="Kuwamura">{{cite journal |last1=Kuwamura |first1=T. |year=1997 |title=Evolution of Female Egg Care in Haremic Triggerfish, ''Rhinecanthus aculeatus'' |journal=[[Ethology (journal)|Ethology]] |volume=103 |issue=12 |pages=1015β1023 |doi=10.1111/j.1439-0310.1997.tb00143.x |bibcode=1997Ethol.103.1015K }}</ref>
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