Editing Old World flycatcher (section)

===Genera===
The family formerly included fewer species. At the time of the publication of the third edition of [[Howard and Moore Complete Checklist of the Birds of the World]] in 2003, the genera ''[[Myophonus]]'', ''[[Alethe (genus)|Alethe]]'', ''[[Brachypteryx]]'' and ''[[Monticola]]'' were included in the thrush family [[Turdidae]].<ref name=h&m3>{{cite book | editor1-last=Dickinson | editor1-first=E.C. | year=2003 | title=The Howard and Moore Complete Checklist of the Birds of the World | edition=3rd | place=London | publisher=Christopher Helm | isbn=978-0-7136-6536-9 }}</ref> Subsequent [[molecular phylogenetic]] studies have shown that the species in these four genera are more closely related to species in Muscicapidae.<ref name=voelker2004>{{ cite journal | last1=Voelker | first1=G. | last2=Spellman | first2=G.M. | year=2004 | title=Nuclear and mitochondrial DNA evidence of polyphyly in the avian superfamily Muscicapoidea | journal=Molecular Phylogenetics and Evolution | volume=30 | issue= 2| pages=386–394 | doi=10.1016/S1055-7903(03)00191-X | pmid=14715230 | bibcode=2004MolPE..30..386V }}</ref><ref name=sangster2010>{{ cite journal | last1=Sangster | first1=G. | last2=Alström | first2=P. | last3=Forsmark | first3=E. | last4=Olsson | first4=U. | year=2010 | title=Multi-locus phylogenetic analysis of Old World chats and flycatchers reveals extensive paraphyly at family, subfamily and genus level (Aves: Muscicapidae) | journal=Molecular Phylogenetics and Evolution | volume=57 | issue=1 | pages=380–392 | doi=10.1016/j.ympev.2010.07.008 | pmid=20656044| bibcode=2010MolPE..57..380S }}</ref> As a consequence, these four genera are now placed here.<ref name=ioc/><ref name=h&m4>{{cite book | editor1-last=Dickinson | editor1-first=E.C. | editor2-last=Christidis | editor2-first=L. | year=2014 | title=The Howard & Moore Complete Checklist of the Birds of the World. | volume=2, Passerines | edition=4th | publisher=Aves Press | place=Eastbourne, U.K. | isbn=978-0-9568611-2-2 | pages=584, 598, 601, 607 }}</ref> In contrast, the genus ''[[Cochoa]]'' which was previously placed in Muscicapidae has been shown to belong in Turdidae.<ref name=voelker2004/><ref name=sangster2010/>

Two large [[molecular phylogenetic]] studies of species within Muscicapidae published in 2010 showed that the genera ''Fraseria'', ''Melaenornis'' and ''Muscicapa'' were [[monophyletic|non-monophyletic]]. The authors were unable to propose revised genera as not all the species were sampled and not all the nodes in their phylogenies were strongly supported.<ref name=sangster2010/><ref>{{ cite journal | last1=Zuccon | first1=D. | last2=Ericson | first2=P.G.P. | year=2010 | title=A multi-gene phylogeny disentangles the chat-flycatcher complex (Aves: Muscicapidae) | journal=Zoologica Scripta | volume=39 | issue=3 | pages=213–224 | doi=10.1111/j.1463-6409.2010.00423.x | s2cid=85963319 }}</ref> A subsequent study published in 2016, that included 37 of the 42 Muscicapini species, confirmed that the genera were non-monophyletic and proposed a reorganised arrangement of the species with several new or resurrected genera.<ref>{{ cite journal | last1=Voelker | first1=G. | last2=Huntley | first2=J.W. | last3=Peñalba | first3=J.V. | last4=Bowie | first4=R.C.K. | year=2016 | title=Resolving taxonomic uncertainty and historical biogeographic patterns in ''Muscicapa'' flycatchers and their allies | journal=Molecular Phylogenetics and Evolution | volume=94 | issue= Pt B| pages=618–625 | doi=10.1016/j.ympev.2015.09.026 | pmid=26475615| doi-access=free | bibcode=2016MolPE..94..618V }}</ref>

[[File:Madagascar magpie-robin (Copsychus albospecularis pica) female.jpg|thumb|Madagascar magpie-robin ''Copsychus albospecularis pica'']]
[[File:Silverbird in Murchison Falls National Park, Uganda.JPG|thumb|right|[[Silverbird (bird)|Silverbird]], monotypic genus ''Empidornis'']]
[[File:Tickell's Blue Flycatcher (Cyornis tickelliae) with feed W IMG 9340.jpg|thumb|[[Cyornis tickelliae|Tickell's blue flycatcher]], genus ''Cyornis'']]
[[File:Amber mountain rock thrush (Monticola sharpei erythronotus) male 2.jpg|thumb|Amber mountain rock thrush ''Monticola sharpei erythronotus'']]
[[File:Stonechat (Saxicola torquatus) male.jpg|thumb|European stonechat ''Saxicola torquatus'']]
[[File:Cyprus wheatear (Oenanthe cypriaca).jpg|thumb|upright|Cyprus wheatear ''Oenanthe cypriaca'']]

The [[International Ornithologists' Union]] recognises 357 species and divides the family into 57 genera. Subdivisions have been proposed by [[George Sangster|Sangster]] ''et al'' (2010).<ref name=ioc/><ref name=sangster2010/> For a complete list of species, see "[[List of Old World flycatcher species]]".

'''Family Muscicapidae'''
* '''Subfamily Muscicapinae''' ([[John Fleming (naturalist)|Fleming]], 1822)
** '''Tribe Copsychini''' (Sundevall, 1872)
*** ''[[Alethe (genus)|Alethe]]'' – alethes
*** ''[[Tychaedon]]'' – scrub robins
*** ''[[Cercotrichas]]'' – scrub robins
*** ''[[Copsychus]]'' – magpie-robins or shamas
** '''Tribe Muscicapini''' (Fleming, 1822)
*** ''[[Agricola (bird)|Agricola]]''
*** ''[[Fraseria]]'' – forest flycatchers
*** ''[[Melaenornis]]'' (7 species)
*** ''[[Herero chat|Namibornis]]'' – single species: Herero chat
*** ''[[Silverbird (bird)|Empidornis]]'' – single species: silverbird
*** ''[[Sigelus]]'' – single species: fiscal flycatcher
*** ''[[Bradornis]]'' (3 species)
*** ''[[Myornis]]'' – single species: Böhm's flycatcher
*** ''[[Artomyias]]'' (2 species)
*** ''[[Humblotia (bird)|Humblotia]]'' – single species: Humblot's flycatcher
*** ''[[Muscicapa]]'' (17 species)
* '''Subfamily Niltavinae''' (Sangster, Alström, Forsmark and Olsson, 2010)<ref name=sangster2016>{{ cite journal | last1=Sangster | first1=G. | last2=Alström | first2=P. | last3=Forsmark | first3=Émile | last4=Olsson | first4=U. | year=2016 | title=Niltavinae, a new taxon of Old World flycatchers (Aves: Muscicapidae) | journal=Zootaxa | volume=4196 | issue=3 | pages=428–429 | doi=10.11646/zootaxa.4196.3.7 | pmid=27988667 | doi-access=free }}</ref>{{efn|The ornithologist Dario Zuccon pointed out that when George Sangster and colleagues erected the name "Niltavinae" for the subfamily, they did not provide a description as required by the [[International Code of Zoological Nomenclature]]. Sangster and colleagues subsequently published a description in 2016.<ref name=sangster2016/><ref name=zuccon2011>{{ cite journal | last=Zuccon | first=D. | year=2011 | title=Taxonomic notes on some Muscicapidae | journal=Bulletin of the British Ornithologists' Club | volume=131 | issue=3 | pages=196–199 | url=https://www.biodiversitylibrary.org/page/50781147 }}</ref>}}
** ''[[Leucoptilon]]'' – single species: white-tailed flycatcher
** ''[[Sholicola]]''<ref>{{cite journal |last1=Robin |first1=V.V. |last2=Vishnudas |first2=C. K. |last3=Gupta |first3=Pooja |last4=Rheindt |first4=Frank E. |last5=Hooper |first5=Daniel M. |last6=Ramakrishnan |first6=Uma |last7=Reddy |first7=Sushma |year=2017 |title=Two new genera of songbirds represent endemic radiations from the Shola Sky Islands of the Western Ghats, India |journal=BMC Evolutionary Biology |volume=17 |issue=1 |pages=31 |doi=10.1186/s12862-017-0882-6 |pmc=5259981 |pmid=28114902 |doi-access=free|bibcode=2017BMCEE..17...31R }}</ref> – sholakilis
** ''[[Niltava]]'' – niltavas
** ''[[Cyanoptila]]'' – flycatchers
** ''[[Eumyias]]'' – blue flycatchers
** ''[[Anthipes]]'' – flycatchers
** ''[[Cyornis]]'' – blue flycatchers
* '''Subfamily Erithacinae''' ([[George Robert Gray|G.R. Gray]], 1846) – African forest robin assemblage{{efn|Dario Zuccon has argued that the correct name for the African forest robins assemblage is Cossyphinae (type genus ''Cossypha'' [[Nicholas Aylward Vigors|Vigors]], 1825) as the name predates Erithacinae (G.R. Gray, 1846).<ref name=zuccon2011/><ref>{{cite journal | last=Vigors | first=Nicholas Aylward | author-link=Nicholas Aylward Vigors | year=1825 | title=''Cossyphina'' | journal=Zoological Journal | volume=2 | page=395 | url=https://www.biodiversitylibrary.org/page/2255532}}</ref>}}
** ''[[Erithacus]]'' – single species: European robin
** ''[[Swynnertonia]]'' – single species: Swynnerton's robin
** ''[[Pogonocichla]]'' – single species: white-starred robin
** ''[[Stiphrornis]]'' – forest robins (3 species)
** ''[[Cossyphicula]]'' – robin-chats
** ''[[Chamaetylas]]'' – (4 species)
** ''[[Cossypha]]'' – robin-chats
** ''[[Cichladusa]]'' – palm thrushes
** ''[[Angola cave chat|Xenocopsychus]]'' – single species: Angola cave chat
** ''[[Dessonornis]]'' – robin-chats
** ''[[Sheppardia]]'' – akalats
* '''Subfamily [[Saxicolinae]]''' ([[Nicholas Aylward Vigors|Vigors]], 1825)
** ''[[Irania (bird)|Irania]]'' – single species: white-throated robin
** ''[[Luscinia]]'' – nightingales and relatives
** ''[[Myiomela]]'' – robins
** ''[[Calliope (genus)|Calliope]]'' – rubythroats
** ''[[Enicurus]]'' – forktails
** ''[[Cinclidium (bird)|Cinclidium]]'' – single species: blue-fronted robin
** ''[[Myophonus]]'' – whistling thrushes
** ''[[Heinrichia]]'' – single species: great shortwing
** ''[[Vauriella]]''
** ''[[Leonardina]]'' – single species: Bagobo babbler
** ''[[Brachypteryx]]'' – shortwings
** ''[[Larvivora]]'' – East and South-East Asian robins
** ''[[Ficedula]]'' – flycatchers
** ''[[Tarsiger]]'' – bush robins and bluetails
** ''[[Heteroxenicus]]'' – single species: Gould's shortwing
** ''[[Phoenicurus]]'' – redstarts
** ''[[Monticola]]'' – rock thrushes
** ''[[Saxicola]]'' – stonechats and chats
** ''[[Campicoloides]]'' – single species: buff-streaked chat
** ''[[Emarginata]]''
** ''[[Pinarochroa]]'' – single species: moorland chat
** ''[[Thamnolaea]]'' – single species: mocking cliff chat
** ''[[Myrmecocichla]]'' – chats
** ''[[Oenanthe (bird)|Oenanthe]]'' – wheatears

The [[cladogram]] below is based on a [[molecular phylogenetic]] study of the family by Min Zhao and collaborators that was published in 2023. Some regions of the phylogenetic tree were not strongly supported by the sequence data.<ref name=zhoa>{{Cite journal | last1=Zhao | first1=M. | last2=Gordon Burleigh | first2=J. | last3=Olsson | first3=U. | last4=Alström | first4=P. | last5=Kimball | first5=R.T. | date=2023 | title=A near-complete and time-calibrated phylogeny of the Old World flycatchers, robins and chats (Aves, Muscicapidae) | journal=Molecular Phylogenetics and Evolution | volume=178 | pages=107646 | doi=10.1016/j.ympev.2022.107646 | doi-access=free | pmid=36265831 | bibcode=2023MolPE.17807646Z }}</ref> Both the genera included and the number of species in each genera are taken from the list of birds maintained by [[Frank Gill (ornithologist)|Frank Gill]], [[Pamela C. Rasmussen]] and David Donsker on behalf of the [[International Ornithological Committee]] (IOC).<ref name=ioc/>

{{Clade | style=font-size:90%;line-height:80%
|label1=Muscicapidae
|1={{Clade
   |label1=Muscicapinae
   |1={{clade
      |label1=Copsychini
      |1={{clade
         |1=''[[Alethe (bird)|Alethe]]'' – alethes (2 species)
         |2={{clade
            |1=''[[Tychaedon]]'' – scrub robins (5 species)
            |2={{clade
               |1=''[[Cercotrichas]]'' – scrub robins (5 species)
               |2=''[[Copsychus]]'' – magpie-robins, sharmas (17 species)
               }}
            }}
         }}
      |label2=Muscicapini
      |2={{clade
         |1=''[[Vauriella goodfellowi]]'' – slaty-backed jungle flycatcher (position uncertain)
         |2={{clade
            |1={{clade
               |1=''[[Agricola (bird)|Agricola]]'' – flycatchers (2 species)
               |2={{clade
                  |1=''[[Fraseria]]'' – flycatchers (8 species)
                  |2={{clade
                     |1=''[[Melaenornis]]'' – flycatchers (7 species)
                     |2={{clade
                        |1=''[[Namibornis]]'' – herero chat
                        |2={{clade
                           |1=''[[Empidornis]]'' – silverbird
                           |2=''[[Sigelus]]'' – fiscal flycatcher
                           }}
                        }}
                     }}
                  }}
               }}
            |2={{clade
               |1={{clade
                  |1=''[[Bradornis]]'' – flycatchers (3 species)
                  |2={{clade
                     |1=''[[Myopornis]]'' – Böhm's flycatcher
                     |2=''[[Artomyias]]'' – flycatchers (2 species)
                     }}
                  }}
               |2=''[[Muscicapa]]'' – flycatchers (17 species) & ''[[Humblotia (bird)|Humblotia]]'' – Humblot's flycatcher
               }}
            }}
         }}
      }}
   |2={{clade
      |label1=Niltavinae
      |1={{clade
         |1=''[[Leucoptilon]]'' – white-tailed flycatcher
         |2={{clade
            |1={{clade
               |1=''[[Sholicola]]'' – blue robins (2 species)
               |2={{clade
                  |1=''[[Niltava]]'' – niltavas (7 species)
                  |2={{clade
                     |1=''[[Cyanoptila]]'' – flycatchers (2 species)
                     |2=''[[Eumyias]]'' – flycatchers (6 species)
                     }}
                  }}
               }}
            |2={{clade
               |1=''[[Anthipes]]'' – flycatchers (2 species)
               |2=''[[Cyornis]]'' – blue flycatchers, jungle flycatchers (36 species) many taxa unsampled
               }}
            }}
         }}
      |2={{clade
         |label1=Cossyphinae
         |1={{clade
            |1=''[[Erithacus]]'' – European robin
            |2={{clade
               |1={{clade
                  |1=''[[Swynnertonia]]'' – Swynnerton's robin
                  |2=''[[Pogonocichla]]'' – white-starred robin
                  }}
               |2={{clade
                  |1={{clade
                     |1=''[[Stiphrornis]]'' – forest robin (position uncertain)
                     |2={{clade
                        |1=''[[Cossyphicula]]'' – robin-chats (2 species)
                        |2={{clade
                           |1=''[[Chamaetylas]]'' – alethes (4 species)
                           |2=''[[Cossypha]]'' - robin-chats (8 species)
                           }}
                        }}
                     }}
                  |2={{clade
                     |1={{clade
                        |1=''[[Cossypha cyanocampter]]'' – blue-shouldered robin-chat (position uncertain)
                        |2={{clade
                           |1=''[[Cichladusa]]'' – palm thrushes (3 species)
                           |2={{clade                         
                              |1=''[[Xenocopsychus]]'' – Angola cave chat
                              |2=''[[Dessonornis]]'' – robin-chats, ground robins (4 species)                                                     
                              }}
                           }}
                        }}
                     |2=''[[Sheppardia]]'' – akalats (11 species)
                     }}
                  }}
               }}
            }}
         |label2=Saxicolinae
          |2={{clade
             |1={{clade
                |1={{clade
                   |1={{clade
                      |1=''[[Irania (bird)|Irania]]'' – white-throated robin
                      |2=''[[Luscinia]]'' – nightingales, redstart, bluethroat (4 species)
                      }}
                   |2=''[[Calliope (genus)|Calliope]]'' – rubythroats (5 species)
                   }}
                |2={{clade
                   |1={{clade
                      |1=''[[Enicurus]]'' – forktails (8 species)
                      |2={{clade
                         |1=''[[Cinclidium (bird)|Cinclidium]]'' – blue-fronted robin
                         |2=''[[Myophonus]]'' – whistling thrushes (9 species)
                         }}
                      }}
                   |2={{clade
                      |1=''[[Myiomela]]'' – robins (3 species)
                      |2={{clade
                         |1={{clade
                            |1={{clade
                               |1=''[[Heinrichia]]'' – great shortwing
                               |2=''[[Vauriella]]'' – jungle flycatchers (3 species) & ''[[Leonardina]]  – ''Bagobo babbler
                               }}
                            |2={{clade
                               |1=''[[Brachypteryx]]'' – shortwings (6 species)
                               |2=''[[Larvivora]]'' – robins (6 species)
                               }}
                            }}
                         |2=''[[Ficedula]]'' – flycatchers (34 species)
                         }}
                      }}
                   }}
                }}
             |2={{clade
                |1=''[[Tarsiger]]'' – bush robins, bluetails (8 species)
                |2={{clade
                   |1=''[[Heteroxenicus]]'' – Gould's shortwing
                   |2={{clade
                      |1=''[[Phoenicurus]]'' – redstarts (14 species)
                      |2={{clade
                         |1=''[[Monticola]]'' – rock thrushes (13 species)
                         |2={{clade
                            |1=''[[Saxicola]]'' – bush chats, stonechats (14 species)
                            |2={{clade
                               |1={{clade
                                  |1=''[[Campicoloides]]'' – buff-streaked chat
                                  |2=''[[Emarginata]]'' – chats (3 species)
                                  }}
                               |2={{clade
                                  |1={{clade
                                     |1=''[[Pinarochroa]]'' – moorland chat
                                     |2=''[[Myrmecocichla]]'' – chats (8 species) & ''[[Thamnolaea]]''  –  mocking cliff chat
                                     }}
                                  |2=''[[Oenanthe (bird)|Oenanthe]]'' – wheatears (32 species)
                                  }}
                              }}
                           }}
                        }}
                     }}
                  }}
               }}
            }}
         }}
      }}
   }}
}}