Editing Pandanus (section)

== Evolution ==
The oldest fossil of the genus is ''Pandanus estellae'', which is known from a [[silicified]] fruit found in Queensland, Australia, dating to the [[Oligocene]] epoch around 32–28 million years ago.<ref>{{Cite journal |last1=Rozefelds |first1=Andrew C. |last2=Rudall |first2=Paula J. |last3=Herne |first3=Matt C. |last4=Milroy |first4=Anita K. |last5=Bridgeman |first5=Joe |date=2022-05-01 |title=A Fossil Syncarpous Fruit from Australia Provides Support for a Gondwanan History for the Screw Pines (Pandanus, Pandanaceae) |journal=International Journal of Plant Sciences |language=en |volume=183 |issue=4 |pages=320–329 |doi=10.1086/719431 |issn=1058-5893 |s2cid=247378720}}</ref> Phylogenetic analyses of the maternal DNA of ''Pandanus'' has shown the genus to be divided into two large groups (Clade I and Clade II), with each of those groups further subdivided into two smaller groups (Subclade Ia, Subclade Ib, Subclade IIa, and Subclade IIb).<ref name=":1">{{Cite journal |last1=Buerki |first1=Sven |last2=Callmander |first2=Martin W. |last3=Devey |first3=Dion S. |last4=Chappell |first4=Lauren |last5=Gallaher |first5=Timothy |last6=Munzinger |first6=Jérôme |last7=Haevermans |first7=Thomas |last8=Forest |first8=Félix |date=2012 |title=Straightening out the screwpines: A first step in understanding phylogenetic relationships within Pandanaceae |journal=Taxon |language=en |volume=61 |issue=5 |pages=1010–1020 |doi=10.1002/tax.615008 |issn=1996-8175}}</ref><ref name=":2">{{Cite journal |last1=Wojahn |first1=John M. A. |last2=Callmander |first2=Martin W. |last3=Buerki |first3=Sven |date=30 January 2025 |title=Pandanus plastomes decoded: When climate mirrors morphology and phylogenetic relationships |journal=American Journal of Botany |language=en |volume= |issue=preprint |pages=e16461 |doi=10.1002/ajb2.16461 |pmid=39887358 |issn=1537-2197}}</ref> The split between Clades I and II was determined to have occurred during the early [[Miocene]], whereas the splits between Subclade Ia and Ib and between Subclade IIa and IIb were determined to have occurred during the [[Middle Miocene Climatic Optimum]].<ref name=":2" />