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==Species== {{Main|Crocus sativus}} ===Description=== [[File:860808-Saffronfarm-01-IMG 7707-2.jpg|alt=Saffron Flowers|thumb|Crocus flowers which yield red saffron stigmas]] [[File:Saffron onions-IMG 7549-2.jpg|alt=Saffron onions|thumb|Corms]] <!-- Deleted image removed: [[File:Saffron-62-1160x773.jpg|alt=Saffron harvest|thumb|Several women in traditional clothes are collecting saffron in one of the farms of [[Torbat-e Heydarieh|Torbat Heydarieh]] city of Khorasan Razavi]] --> The domesticated saffron crocus, ''Crocus sativus'', is an autumn-[[flowering plant|flowering]] [[perennial plant]] unknown in the wild. It possibly descends from the eastern Mediterranean autumn-flowering ''[[Crocus cartwrightianus]]'' which is also known as "wild saffron"{{Sfn|Kafi et al.|2006|p=24}} and is native to [[mainland Greece]], [[Euboea]], [[Crete]], [[Skyros]] and some islands of the [[Cyclades]].<ref name="Atticasaffron">{{cite journal |last1=Jacobsen |first1=Niels |last2=Ørgaard |first2=Marian |title=Crocus cartwrightianus on the Attica Peninsula |journal=ISHS Acta Horticulturae |date=2004 |volume=650 |issue=6 |pages=65–69 |doi=10.17660/ActaHortic.2004.650.6 |url=https://safranerio.fr/wp-content/uploads/2017/04/650_6.pdf |access-date=6 April 2024 |ref=Cartwright}}</ref> The similar species ''[[Crocus thomasii|C. thomasii]]'' and ''[[Crocus pallasii|C. pallasii]]'' were considered as other possible ancestors.{{Sfn|Grilli Caiola|2003|p=1}}{{Sfn|Negbi|1999|p=28}} As a genetically monomorphic clone incapable of seed production, it was slowly propagated by humans throughout much of [[Eurasia]].{{Sfn|Rubio-Moraga|Castillo-López|Gómez-Gómez|Ahrazem|2009}} Various origins have been suggested for saffron, including [[Iran]],<ref name="Ghorbani et al.">{{Cite book |first1=R. |last1=Ghorbani |first2=A. |last2=Koocheki |s2cid=28214061 |chapter=Sustainable Cultivation of Saffron in Iran |chapter-url=https://books.google.com/books?id=cdksDwAAQBAJ&pg=PA170 |title=Sustainable Agriculture Reviews |editor-last=Lichtfouse |editor-first=Eric |year=2017 |publisher=Springer |isbn=978-3-319-58679-3 |doi=10.1007/978-3-319-58679-3 |pages=170–171|url=https://hal.inrae.fr/hal-02801642/file/Sustainable%20Agriculture%20Reviews%2014%20-%20Front%20Matter_1.pdf }}</ref> [[Greece]],<ref name="Gresta et al.">{{Cite journal|first1=F. |last1=Gresta |first2=G. M. |last2=Lombardo |first3=L. |last3=Siracusa |first4=G. |last4=Ruberto |s2cid=44054590 |year=2008 |title=Saffron, an alternative crop for sustainable agricultural systems. A review |url=https://hal.archives-ouvertes.fr/hal-00886393/document |journal=Agronomy for Sustainable Development |volume=28 |issue=1 |pages=95–112 |doi=10.1051/agro:2007030|bibcode=2008AgSD...28...95G }}</ref> [[Mesopotamia]].<ref name="Ghorbani et al." /> and [[Kashmir]].<ref name="auto">{{Cite book |url=https://www.worldcat.org/oclc/1140113593 |title=Saffron : science, technology and health |date=2020 |others=A. Koocheki, Mohammad Khajeh-Hosseini |isbn=978-0-12-818740-1 |location=Duxford, United Kingdom |oclc=1140113593 |access-date=28 March 2022 |archive-date=19 January 2023 |archive-url=https://web.archive.org/web/20230119180710/https://www.worldcat.org/title/1140113593 |url-status=live }}</ref> It is a sterile [[polyploidy|triploid]] form, which means that three homologous sets of [[chromosome]]s make up each specimen's genetic complement; ''C. sativus'' bears eight chromosomal bodies per set, making for 24 in total.{{Sfn|Kafi et al.|2006|p=23}} Being sterile, the purple flowers of ''C. sativus'' fail to produce viable seeds; reproduction hinges on human assistance: clusters of [[corm]]s, underground, bulb-like, starch-storing organs, must be dug up, divided, and replanted. A corm survives for one season, producing via vegetative division up to ten "cormlets" that can grow into new plants in the next season.{{Sfn|Deo|2003|p=1}} The compact corms are small, brown globules that can measure as large as {{Convert|5|cm|in|0|abbr=on}} in diameter, have a flat base, and are shrouded in a dense mat of parallel fibres; this coat is referred to as the "corm tunic". Corms also bear vertical fibres, thin and net-like, that grow up to {{Convert|5|cm|in|0|abbr=on}} above the plant's neck.{{Sfn|Kafi et al.|2006|p=23}} The plant sprouts 5–11 white and non-[[photosynthesis|photosynthetic]] leaves known as [[cataphyll]]s. These membrane-like structures cover and protect 5 to 11 true leaves as they bud and develop on the crocus flower. The latter are thin, straight, and blade-like green foliage leaves, which are {{Convert|1|–|3|mm|in|frac=32|abbr=on}}, in diameter, which either expand after the flowers have opened ("hysteranthous") or do so simultaneously with their blooming ("synanthous"). ''C. sativus'' cataphylls are suspected by some to manifest prior to blooming when the plant is irrigated relatively early in the growing season. Its floral axes, or flower-bearing structures, bear [[bract]]eoles, or specialised leaves, that sprout from the flower stems; the latter are known as [[pedicel (botany)|pedicels]].{{Sfn|Kafi et al.|2006|p=23}} After [[Aestivation (botany)|aestivating]] in spring, the plant sends up its true leaves, each up to {{Convert|40|cm|in|0|abbr=on}} in length. Only in October, after most other flowering plants have released their seeds, do its brilliantly hued flowers develop; they range from a light pastel shade of lilac to a darker and more striated mauve.{{Sfn|Willard|2002|p=3}} The flowers possess a sweet, honey-like fragrance. Upon flowering, the plants are {{Convert|20|–|30|cm|in|0|abbr=on}} in height and bear up to four flowers. A three-pronged [[style (botany)|style]] {{Convert|25|–|30|mm|in|frac=16|abbr=on}} in length, emerges from each flower. Each prong terminates with a vivid crimson [[stigma (botany)|stigma]], which is the distal end of a [[carpel]].{{Sfn|Deo|2003|p=1}}{{Sfn|Kafi et al.|2006|p=23}} ===Cultivation=== <!-- Deleted image removed: [[File:Iran-MA-20221554-1160x1740.jpg|alt=cleaning Saffron |thumb|An Iranian woman is cleaning saffron.]] --> The saffron crocus, unknown in the wild, probably descends from ''[[Crocus cartwrightianus]]''. It is a [[polyploid|triploid]] that is "self-incompatible" and male sterile; it undergoes aberrant [[meiosis]] and is hence incapable of independent sexual reproduction—all propagation is by [[vegetative reproduction|vegetative multiplication]] via manual "divide-and-set" of a starter clone or by interspecific hybridisation.{{Sfn|Negbi|1999|p=30–31}}{{Sfn|Grilli Caiola|2003|p=1}} ''Crocus sativus'' thrives in the Mediterranean [[Maquis shrubland|maquis]], an ecotype superficially resembling the North American [[chaparral]], and similar climates where hot and dry summer breezes sweep semi-arid lands. It can nonetheless survive cold winters, tolerating frosts as low as {{Convert|−10|C|0}} and short periods of snow cover.{{Sfn|Deo|2003|p=1}}{{Sfn|Willard|2002|pp=2–3}} Some reports suggest saffron can tolerate an air temperature range from −22 to 40 °C.<ref>{{Citation |last=Rezvani-Moghaddam |first=Parviz |title=Chapter 8. Ecophysiology of saffron |date=2020-01-01 |url=https://www.sciencedirect.com/science/article/pii/B9780128186381000083 |work=Saffron |pages=119–137 |editor-last=Koocheki |editor-first=Alireza |access-date=2023-02-22 |series=Woodhead Publishing Series in Food Science, Technology and Nutrition |publisher=Woodhead Publishing |language=en |isbn=978-0-12-818638-1 |editor2-last=Khajeh-Hosseini |editor2-first=Mohammad}}</ref> Irrigation is required if grown outside of moist environments such as Kashmir, where annual rainfall averages {{Convert|1000|–|1500|mm|in|sigfig=1|abbr=on}}; saffron-growing regions in Greece ({{Convert|500|mm|in|abbr=on|disp=or}} annually) and Spain ({{Convert|400|mm|in|abbr=on|disp=or}}) are far drier than the main cultivating Iranian regions. What makes this possible is the timing of the local wet seasons; generous spring rains and drier summers are optimal. Rain immediately preceding flowering boosts saffron yields; rainy or cold weather during flowering promotes disease and reduces yields. Persistently damp and hot conditions harm the crops,{{Sfn|Deo|2003|p=2}} and rabbits, rats, and birds cause damage by digging up corms. [[Nematode]]s, leaf [[rust (fungus)|rusts]], and corm rot pose other threats. Yet ''[[Bacillus subtilis]]'' inoculation may provide some benefit to growers by speeding corm growth and increasing stigma biomass yield.{{Sfn|Sharaf-Eldin|Elkholy|Fernández|Junge|2008}} The plants fare poorly in shady conditions; they grow best in full sunlight. Fields that slope towards the sunlight are optimal (i.e., south-sloping in the Northern Hemisphere). Planting is mostly done in June in the Northern Hemisphere, where corms are lodged {{Convert|7|–|15|cm|in|0|abbr=on}} deep; its roots, stems, and leaves can develop between October and February.{{Sfn|Kafi et al.|2006|p=23}} Planting depth and corm spacing, in concert with climate, are critical factors in determining yields. Mother corms planted deeper yield higher-quality saffron, though they form fewer flower buds and daughter corms. Italian growers optimise thread yield by planting {{Convert|15|cm|in|0|abbr=on}} deep and in rows {{Convert|2|–|3|cm|in|frac=4|abbr=on}} apart; depths of {{Convert|8|–|10|cm|in|0|abbr=on}} optimise flower and corm production. Greek, Moroccan, and Spanish growers employ distinct depths and spacings that suit their locales. ''C. sativus'' prefers friable, loose, low-density, well-watered, and well-drained clay-[[calcareous]] soils with high organic content. Traditional raised beds promote good drainage. Soil organic content was historically boosted via application of some {{Convert|20|–|30|t/ha|ST/acre|0|abbr=off}} of manure. Afterwards, and with no further manure application, corms were planted.{{Sfn|Deo|2003|p=3}} After a period of dormancy through the summer, the corms send up their narrow leaves and begin to bud in early autumn. Only in mid-autumn do they flower. Harvests are by necessity a speedy affair: after blossoming at dawn, flowers quickly wilt as the day passes.{{Sfn|Willard|2002|pp=3–4}} All plants bloom within a window of one or two weeks.{{Sfn|Willard|2002|p=4}} Stigmas are dried quickly upon extraction and (preferably) sealed in airtight containers.{{Sfn|Negbi|1999|p=8}} ===Harvesting=== [[File:Saffron-sargol-zafaranic.jpg|thumb|Sargol saffron, the strongest Iranian grade]] The high retail value of saffron is maintained on world markets because of labour-intensive harvesting methods, which require some {{convert|200000|/lb|/kg|order=flip|disp=preunit|stigmas|hand-picked saffron stigmas }} – equivalently, {{convert|70000|/lb|/kg|order=flip|disp=preunit|flowers|crocus flowers }}.<ref name=hooker/><ref name="cnn">{{cite news |last=Monks |first=Keiron |date=3 September 2015 |title=Iran's homegrown treasure: the spice that costs more than gold |url=https://edition.cnn.com/2015/06/03/middleeast/iran-saffron-red-gold/ |newspaper=CNN |access-date=22 January 2016}}</ref>{{Sfn|Hill|2004|p=273}}{{Sfn|Rau|1969|p=35}} Forty hours of labour are needed to pick 150,000 flowers.<ref>{{cite news|last=Lak |first=Daniel |date=11 November 1998 |title=Kashmiris pin hopes on saffron |work=BBC News |url=http://news.bbc.co.uk/1/hi/world/south_asia/212491.stm |access-date=11 September 2011}}</ref> One freshly picked crocus flower yields on average 30 mg of fresh saffron or 7 mg dried; roughly 150 flowers yield {{Convert|1|g|oz|frac=32|abbr=on}} of dry saffron threads; to produce {{Convert|12|g|oz|frac=16|abbr=on}} of dried saffron, {{Convert|1|lb|g|order=flip|sigfig=2|abbr=on}} of flowers are needed; the yield of dried spice from fresh saffron is only {{convert|0.2|oz/lb|g/kg|order=flip|abbr=on}}.{{Sfn|Deo|2003|p=3}}
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