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===Lifespan=== Turtles can live long lives. The oldest living turtle and land animal is said to be a [[Seychelles giant tortoise]] named [[Jonathan (tortoise)|Jonathan]], who turned 187 in 2019.<ref>{{cite web |url=https://www.guinnessworldrecords.com/news/2019/2/introducing-jonathan-the-worlds-oldest-animal-on-land-561882 |title=Introducing Jonathan, the World's Oldest Animal on Land at 187 Years Old |last=Millward |first=Adam |publisher=[[Guinness World Records]] |date=February 27, 2019 |access-date=July 14, 2020 |archive-date=August 9, 2020 |archive-url=https://web.archive.org/web/20200809153110/https://www.guinnessworldrecords.com/news/2019/2/introducing-jonathan-the-worlds-oldest-animal-on-land-561882 |url-status=live }}</ref> A [[Galápagos tortoise]] named [[Harriet (tortoise)|Harriet]] was collected by [[Charles Darwin]] in 1835; it died in 2006, having lived for at least 176 years. Most wild turtles do not reach that age. Turtles keep growing new scutes under the previous scutes every year, allowing researchers to estimate how long they have lived.{{sfn|Franklin|2011|pp=44–45}} They also [[negligible senescence|age slowly]].<ref name="oldage">{{cite journal |last1=Warner |first1=Daniel A. |last2=Miller |first2=David A. W. |last3=Bronikowski |first3=Anne M. |last4=Janzen |first4=Fredric J. |year=2016 |title=Decades of Field Data Reveal that Turtles Senesce in the Wild |journal=PNAS |volume=113 |issue=23 |pages=6502–6507 |doi=10.1073/pnas.1600035113 |pmid=27140634 |pmc=4988574 |bibcode=2016PNAS..113.6502W |doi-access=free }}</ref> The survival rate for adult turtles can reach 99% per year.<ref name=Firefly/>
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