Editing Quantitative genetics (section)

==== Total and partitioned dispersed quasi-dominance variances====
The ''total genic variance'' is of intrinsic interest in its own right. But, prior to the refinements by Gordon,<ref name="Gordon 2003"/> it had had another important use as well. There had been no extant estimators for the "dispersed" quasi-dominance. This had been estimated as the difference between Sewall Wright's ''inbred genotypic variance'' <ref name="Wright 1951"/> and the total "dispersed" genic variance [see the previous sub-section]. An anomaly appeared, however, because the ''total quasi-dominance variance'' appeared to increase early in inbreeding despite the decline in heterozygosity.<ref name="Falconer 1996"/> {{rp|128}} {{rp|266}}

The refinements in the previous sub-section corrected this anomaly.<ref name="Gordon 2003"/> At the same time, a direct solution for the ''total quasi-dominance variance'' was obtained, thus avoiding the need for the "subtraction" method of previous times. Furthermore, direct solutions for the ''amongst-line'' and ''within-line'' partitions of the ''quasi-dominance variance'' were obtained also, for the first time. [These have been presented in the section "Dispersion and the genotypic variance".]