Editing Concertina movement (section)

=== Arboreal ===

Arboreal concertina locomotion is employed on bare branches on trees, when secondary branches are not available (when they are, snakes perform lateral undulation, using these branches as contact points).<ref name = "Astley">{{cite journal
 | doi         = 10.1242/jeb.009050 
 | doi-access  = free
 | first1      = H.C.
 | first2      = B.C.
 | issue       = Pt 21
 | journal     = Journal of Experimental Biology
 | last1       = Astley
 | last2       = Jayne
 | pages       = 3862–3872
 | pmid        = 17951427
 | title       = Effects of perch diameter and incline on the kinematics, performance and modes of arboreal locomotion of corn snakes (''Elaphe guttata'') 
 | volume      = 210
 | year        = 2007 
}}</ref> In this mode of concertina locomotion, snakes likely grip the branch by ventrally flexing the body at the points where the alternation bends cross the perch, with the bends themselves sometimes extending beyond the edge of the branch. The snake extends the anterior portion of its body, but as it does so, the body follows a constant path (like lateral undulation, but unlike tunnel concertina locomotion). It then forms anterior grips and pulls the body forward, again demonstrating the 'path following' characteristic. Unlike tunnel concertina locomotion, this mode avoids any obstacle which falls between the bends of the snake's body.<ref name="Astley" /> However, it is exceptionally slow, with snakes rarely moving faster than 2% of their length per second.<ref name="Astley" />

Some snakes, such as the brown tree snake, use a "lasso-climbing" or "lasso locomotion" technique to climb vertically by wrapping around the vertical object, including poles intended to stop predators.<ref>{{cite news
 | access-date = 11 January 2021
 | date        = 11 January 2021
 | first1      = Theresa
 | last1       = Machemer
 | title       = Invasive Brown Tree Snakes Stun Scientists With Amazing New Climbing Tactic
 | url         = https://smithsonianmag.com/science-nature/invasive-brown-tree-snakes-stun-scientists-amazing-new-climbing-tactic-180976728
 | work        = [[Smithsonian (magazine)|Smithsonian]]
}}</ref><ref>{{cite news
 | access-date = 2 August 2023
 | date        = 11 January 2021
 | first1      = Julie A.
 | first2      = Thomas F.
 | first3      = Martin
 | first4      = Bruce C.
 | last1       = Savidge
 | last2       = Seibert
 | last3       = Kastner
 | last4       = Jayne
 | title       = Lasso locomotion expands the climbing repertoire of snakes
 | url         = https://doi.org/10.1016/j.cub.2020.11.050
 | work        = Current Biology
}}</ref>