Editing Cotinga (section)

==Taxonomy and systematics==
The family Cotingidae was introduced by French naturalist [[Charles Lucien Bonaparte]] in 1849.<ref>{{cite book | last=Bock | first=Walter J. | year=1994 | title=History and Nomenclature of Avian Family-Group Names | series=Bulletin of the American Museum of Natural History | volume= 222 | publisher=American Museum of Natural History | place=New York | page=149 | hdl=2246/830 }}<!--Link allows download of the 48MB pdf--></ref><!--Bock gives Bonaparte's title as "Conspectus systematis ornithologiae. Editio altera reformata additis synonymis Grayanis" 1849. This doesn't appear to be available on the web. It is discussed at https://www.birdforum.net/showthread.php?t=316935&page=3 There is what appears to be an 1851 reprint: Bonaparte CL. 1851. Conspectus systematis ornithologiae. Editio altera reformata additis synonymis Grayanis. Nuov. Ann. Sci. Nat. (ser. 3) 3:480-482; 4:196-206. available at: https://biodiversitylibrary.org/page/9363469 --> According to the [[International Ornithological Committee]], as of July 2021, the family contains 66 species divided into 24 genera.<ref name=ioc>{{cite web| editor1-last=Gill | editor1-first=Frank | editor1-link=Frank Gill (ornithologist) | editor2-last=Donsker | editor2-first=David | editor3-last=Rasmussen | editor3-first=Pamela | editor3-link=Pamela Rasmussen | date=July 2021 | title=Cotingas, manakins, tityras, becards | work=IOC World Bird List Version 11.2 | url=http://www.worldbirdnames.org/bow/cotingas/ | publisher=International Ornithologists' Union | access-date=7 December 2021 }}</ref>

A 2014 [[molecular phylogenetic]] study of the cotingas by Jacob Berv and [[Richard Prum]] found that the genera formed five [[monophyletic]] clades and they proposed that the family could be divided into five subfamilies.<ref name=berv>{{ cite journal | last1=Berv | first1=J.S. | last2=Prum | first2=R.O. | author2-link=Richard Prum  | year=2014 | title=A comprehensive multilocus phylogeny of the Neotropical cotingas (Cotingidae, Aves) with a comparative evolutionary analysis of breeding system and plumage dimorphism and a revised phylogenetic classification | journal=Molecular Phylogenetics and Evolution | volume=81 | pages=120–136 | doi=10.1016/j.ympev.2014.09.001 | pmid=25234241 }}</ref> The following cladogram is based on a molecular phylogenetic study of the [[suboscine]]s by Michael Harvey and collaborators published in 2020.<ref>{{Cite journal | last1=Harvey | first1=M.G. | last2=Bravo | first2=G.A. | last3=Claramunt | first3=S. | last4=Cuervo | first4=A.M. | last5=Derryberry | first5=G.E. | last6=Battilana | first6=J. | last7=Seeholzer | first7=G.F. | last8=McKay | first8=J.S. | last9=O’Meara | first9=B.C. | last10=Faircloth | first10=B.C. | last11=Edwards | first11=S.V. | last12=Pérez-Emán | first12=J. | last13=Moyle | first13=R.G. | last14=Sheldon | first14=F.H. | last15=Aleixo | first15=A. | last16=Smith | first16=B.T. | last17=Chesser | first17=R.T. | last18=Silveira | first18=L.F. | last19=Cracraft | first19=J. | last20=Brumfield | first20=R.T. | last21=Derryberry | first21=E.P. | date=2020 | title=The evolution of a tropical biodiversity hotspot | journal=Science | volume=370 | issue=6522 | pages=1343–1348 | doi=10.1126/science.aaz6970 | pmid=33303617 | bibcode=2020Sci...370.1343H | hdl=10138/329703 | s2cid=228084618 | hdl-access=free }}  A high resolution version of the phylogenetic tree in Figure 1 is available from the first author's website [http://www.harveybirdlab.org/docs/Harveyetal2020_Fig1_tree_HiRes.pdf here].</ref>
{{Clade| style=font-size:80%;line-height:100%
|label1=Cotingidae
|1={{clade
   |label1=Pipreolinae
   |1={{clade
      |1=''[[Ampelioides]]'' – scaled fruiteater
      |2=''[[Pipreola]]'' – fruiteaters (11 species)
      }}
   |2={{clade
      |label1=Rupicolinae
      |1={{clade
         |1={{clade
            |1=''[[Snowornis]]'' – pihas (2 species)
            |2=''[[Carpornis]]'' – berryeaters (2 species)
            }}
         |2={{clade
            |1=''[[Phoenicircus]]'' – cotingas (2 species)
            |2=''[[Cock-of-the-rock|Rupicola]]'' – cock-of-the-rocks (2 species)
            }}
         }}
      |2={{clade
         |label1=Phytotominae
         |1={{clade
            |1=''[[Zaratornis]]'' – white-cheeked cotinga
            |2={{clade
               |1=''[[Phytotoma]]'' – plantcutters (3 species)
               |2={{clade
                  |1=''[[Phibalura]]'' – cotingas (1 species)
                  |2={{clade
                     |1=''[[Doliornis]]'' – cotingas (2 species)
                     |2=''[[Ampelion]]'' – cotingas (2 species)
                     }}
                  }}
               }}
            }}
         |2={{clade
            |label1=Cephalopterinae
            |1={{clade
               |1=''[[Haematoderus]]'' – crimson fruitcrow
               |2={{clade
                  |1=''[[Querula]]'' –  purple-throated fruitcrow
                  |2={{clade
                     |1=''[[Pyroderus]]'' – red-ruffed fruitcrow
                     |2={{clade
                        |1=''[[Perissocephalus]]'' – capuchinbird 
                        |2=''[[Cephalopterus]]'' – umbrellabirds (3 species)
                        }}
                     }}
                  }}
               }}
            |label2=Cotinginae
            |2={{clade
               |1=''[[Lipaugus]]'' – pihas and cotingas (9 species)
               |2={{clade
                  |1=''[[Procnias]]'' – bellbirds (4 species)
                  |2={{clade
                     |1=''[[Cotinga (genus)|Cotinga]]'' – cotingas (7 species)
                     |2={{clade
                        |1=''[[Porphyrolaema]]'' – purple-throated cotinga
                        |2={{clade
                           |1={{clade
                              |1=''[[Conioptilon]]'' – black-faced cotinga
                              |2=''[[Gymnoderus]]'' – bare-necked fruitcrow
                              }}
                           |2={{clade
                              |1=''[[Xipholena]]'' – cotingas (3 species)
                              |2=''[[Carpodectes]]'' – cotingas (3 species)
                              }}
                           }}
                        }}
                     }}
                  }}
               }}
            }}
         }}
      }}
   }}
}}
The genus ''Tijuca'' was found to be embedded in ''Lipaugus'', a position that was confirmed by a more detailed 2020 study.<ref>{{Cite journal | last1=Settlecowski | first1=A.E. | last2=Cuervo | first2=A.M. | last3=Tello | first3=J.G. | last4=Harvey | first4=M.G. | last5=Brumfield | first5=R.T. | last6=Derryberry | first6=E.P. | date=2020 | title=Investigating the utility of traditional and genomic multilocus datasets to resolve relationships in ''Lipaugus'' and ''Tijuca'' (Cotingidae) | journal=Molecular Phylogenetics and Evolution  | volume=147 | pages=106779 | doi=10.1016/j.ympev.2020.106779| pmid=32135309 | s2cid=212568712 }}</ref>

{| class="wikitable"
|-
! Image !! Genus  !! Living species
|-
|[[File:Ampelioides tschudii (Frutero escamado) (24) (14148007464).jpg|175px]] || ''[[Ampelioides]]'' {{small|Verreaux, 1867}} || 
* [[Scaled fruiteater]], ''Ampelioides tschudii''
|-
|[[File:Pipreola frontalis squamipectus.jpg|175px]] || ''[[Pipreola]]'' {{small|Swainson, 1838}}   ||
* [[Fiery-throated fruiteater]], ''Pipreola chlorolepidota''
* [[Scarlet-breasted fruiteater]], ''Pipreola frontalis''
* [[Handsome fruiteater]], ''Pipreola formosa''
* [[Red-banded fruiteater]], ''Pipreola whitelyi'' 
* [[Black-chested fruiteater]], ''Pipreola lubomirskii''
* [[Orange-breasted fruiteater]], ''Pipreola jucunda''
* [[Masked fruiteater]], ''Pipreola pulchra''
* [[Golden-breasted fruiteater]], ''Pipreola aureopectus''
* [[Barred fruiteater]], ''Pipreola arcuata''
* [[Band-tailed fruiteater]], ''Pipreola intermedia''
* [[Green-and-black fruiteater]], ''Pipreola riefferii''
|-
|[[File:Olivaceous Piha - Colombia S4E3368 (16835513815).jpg|175px]] || ''[[Snowornis]]'' {{small|Prum, 2001}}   || 
* [[Grey-tailed piha]], ''Snowornis subalaris''
* [[Olivaceous piha]], ''Snowornis cryptolophus''
|-
|[[File:Hooded berryeater (Carpornis cucullata).jpg|175px]] || ''[[Carpornis]]'' {{small|G.R. Gray, 1846}}   || 
* [[Hooded berryeater]], ''Carpornis cucullata''
* [[Black-headed berryeater]], ''Carpornis melanocephala''
|-
|[[File:Rupicola peruviana (male) -San Diego Zoo-8a.jpg|175px]] ||''[[Cock-of-the-rock|Rupicola]]'' {{small|Brisson, 1760}}  || 
* [[Andean cock-of-the-rock]], ''Rupicola peruvianus''
* [[Guianan cock-of-the-rock]], ''Rupicola rupicola''
|-
|[[File:Phoenicircus carnifex - Guianan red cotinga (male) 01.JPG|175px]] || ''[[Phoenicircus]]'' {{small|Swainson, 1832}}   || 
* [[Guianan red cotinga]], ''Phoenicircus carnifex''
* [[Black-necked red cotinga]], ''Phoenicircus nigricollis''
|-
|[[File:Zaratornis stresemanni - White-cheeked Cotinga.jpg|175px]] || ''[[Zaratornis]]'' {{small|Koepcke, 1954}}   || 
* [[White-cheeked cotinga]], ''Zaratornis stresemanni''
|-
|[[File:White-tipped Plantcutter (Phytotoma rutila).jpg|175px]] || ''[[Phytotoma]]'' {{small|Molina, 1782}}   || 
* [[Rufous-tailed plantcutter]] or Chilean plantcutter, ''Phytotoma rara''
* [[Peruvian plantcutter]], ''Phytotoma raimondii''
* [[White-tipped plantcutter]] or reddish plantcutter, ''Phytotoma rutila''
|-
|[[File:Swallow-tailed Cotinga (Phibalura flavirostris).jpeg|175px]] ||''[[Phibalura]]'' {{small|Vieillot, 1816}}   || 
* [[Swallow-tailed cotinga]], ''Phibalura flavirostris''
|-
|[[File:Doliornis remseni 262570674.jpg|175px]]|| ''[[Doliornis]]'' {{small|Taczanowski, 1874}}   || 
* [[Bay-vented cotinga]], ''Doliornis sclateri''
* [[Chestnut-bellied cotinga]], ''Doliornis remseni''
|-
|[[File:Red-crested Cotinga - EcuadorDSCN2925.jpg|175px]] ||''[[Ampelion]]'' {{small|Tschudi, 1845}}   || 
* [[Red-crested cotinga]], ''Ampelion rubrocristatus''
* [[Chestnut-crested cotinga]], ''Ampelion rufaxilla''
|-
| || ''[[Haematoderus]]'' {{small|Bonaparte, 1854}}   || 
* [[Crimson fruitcrow]], ''Haematoderus militaris''
|-
|[[File:Purple-throated Fruitcrow RWD.jpg|175px]] ||''[[Querula]]'' {{small|Vieillot, 1816}}  || 
* [[Purple-throated fruitcrow]], ''Querula purpurata''
|-
|[[File:Pavó no Parque Estaudal Intervales.jpg|175px]] || ''[[Pyroderus]]'' {{small|G.R. Gray, 1840}}  ||  
* [[Red-ruffed fruitcrow]], ''Pyroderus scutatus''
|-
|[[File:Cephalopterus glabricollis.jpg|175px]] || ''[[Cephalopterus]]'' {{small|E. Geoffroy Saint-Hilaire, 1809}}   ||
*[[Bare-necked umbrellabird]], ''Cephalopterus glabricollis'' 
*[[Long-wattled umbrellabird]], ''Cephalopterus penduliger''
*[[Amazonian umbrellabird]], ''Cephalopterus ornatus''
|-
|[[File:Capuchinbird - Perissocephalus tricolor.jpg|175px]] || ''[[Perissocephalus]]'' {{small|Oberholser, 1899}}  ||  
* [[Capuchinbird]], ''Perissocephalus tricolor''
|-
|[[File:TROPEIRO-DA-SERRA (Lipaugus lanioides) (13868684655).jpg|175px]] || ''[[Lipaugus]]'' {{small|F. Boie, 1828}}  || 
* [[Rufous piha]], ''Lipaugus unirufus''
* [[Cinnamon-vented piha]], ''Lipaugus lanioides''
* [[Rose-collared piha]], ''Lipaugus streptophorus''
* [[Screaming piha]], ''Lipaugus vociferans''
* [[Dusky piha]], ''Lipaugus fuscocinereus''
* [[Scimitar-winged piha]], ''Lipaugus uropygialis''
* [[Chestnut-capped piha]], ''Lipaugus weberi''
* [[Black-and-gold cotinga]], ''Lipaugus ater''
* [[Grey-winged cotinga]], ''Lipaugus conditus''
|-
|[[File:Procnias averano.jpg|175px]] ||''[[Procnias]]'' {{small|Illiger, 1811}}   || 
* [[White bellbird]], ''Procnias albus''
* [[Three-wattled bellbird]], ''Procnias tricarunculatus''
* [[Bare-throated bellbird]], ''Procnias nudicollis''
* [[Bearded bellbird]], ''Procnias averano''
|-
|[[File:SpangledCotinga.jpg|175px]] || ''[[Cotinga (genus)|Cotinga]]'' {{small|Brisson, 1760}}   || 
* [[Plum-throated cotinga]], ''Cotinga maynana''
* [[Spangled cotinga]], ''Cotinga cayana''
* [[Lovely cotinga]], ''Cotinga amabilis''
* [[Blue cotinga]], ''Cotinga nattererii''
* [[Turquoise cotinga]], ''Cotinga ridgwayi''
* [[Banded cotinga]], ''Cotinga maculata''
* [[Purple-breasted cotinga]], ''Cotinga cotinga''
|-
|[[File:Porphyrolaema porphyrolaema - Purple-throated cotinga (female).JPG|175px]] || ''[[Porphyrolaema]]'' {{small|Bonaparte, 1854}}   || 
* [[Purple-throated cotinga]], ''Porphyrolaema porphyrolaema''
|-
|[[File:Conioptilon mcilhennyi - Black-faced cotinga.jpg|175px]] || ''[[Conioptilon]]'' {{small|Lowery & O'Neill, 1966}}   || 
* [[Black-faced cotinga]], ''Conioptilon mcilhennyi''
|-
|[[File:Gymnoderus foetidus - Bare-necked Fruitcrow.JPG|175px]] || ''[[Gymnoderus]]'' {{small|E. Geoffroy Saint-Hilaire, 1809}}  ||  
* [[Bare-necked fruitcrow]], ''Gymnoderus foetidus''
|-
|[[File:Xipholena punicea -Miami Metrozoo -male-8a.jpg|175px]] || ''[[Xipholena]]'' {{small|Gloger, 1841}}    || 
* [[Pompadour cotinga]], ''Xipholena punicea''
* [[White-tailed cotinga]], ''Xipholena lamellipennis''
* [[White-winged cotinga]], ''Xipholena atropurpurea''
|-
|[[File:Snowy Cotinga - Sarapiqui - Costa Rica S4E1046 (26584977742).jpg|175px]] || ''[[Carpodectes]]'' {{small|Salvin, 1865}}  || 
* [[Black-tipped cotinga]], ''Carpodectes hopkei''
* [[Yellow-billed cotinga]], ''Carpodectes antoniae''
* [[Snowy cotinga]], ''Carpodectes nitidus''
|-
|}

A number of species previously placed in this family are now placed in the family [[Tityridae]] (genera ''[[Laniisoma]]'', ''[[Laniocera]]'' and ''[[Iodopleura]]'')<ref>Remsen, J. V. Jr., C. D. Cadena, A. Jaramillo, M. Nores, J. F. Pacheco, M. B. Robbins, T. S. Schulenberg, F. G. Stiles, D. F. Stotz, & K. J. Zimmer. 2007. [http://www.museum.lsu.edu/~Remsen/SACCBaseline.html ''A classification of the bird species of South America.''] {{webarchive |url=https://web.archive.org/web/20090302073659/http://www.museum.lsu.edu/~Remsen/SACCBaseline.html |date=March 2, 2009 }} American Ornithologists' Union. Accessed 12 December 2007.</ref>

{{See also|List of cotinga species}}