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Craniate
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==Classification== Below is a [[phylogenetic tree]] of the phylum [[Chordata]]. Lines show probable evolutionary relationships, including extinct taxa, which are denoted with a [[Dagger (typography)|dagger]], †. Some groups in this tree ([[lancelets]] and [[tunicates]]) are invertebrates. The positions (relationships) of the lancelet, tunicate, and craniate clades are as reported.<ref>{{Cite journal | title=The amphioxus genome and the evolution of the chordate karyotype | author=Putnam, N. H. | author2=Butts, T. | author3=Ferrier, D. E. K. | author4=Furlong, R. F. | author5=Hellsten, U. | author6=Kawashima, T. | author7=Robinson-Rechavi, M. | author8=Shoguchi, E. | author9=Terry, A. | author10=Yu, J. K. | author11=Benito-Gutiérrez, E. L. | author12=Dubchak, I. | author13=Garcia-Fernàndez, J. | author14=Gibson-Brown, J. J. | author15=Grigoriev, I. V. | author16=Horton, A. C. | author17=De Jong, P. J. | author18=Jurka, J. | author19=Kapitonov, V. V. | author20=Kohara, Y. | author21=Kuroki, Y. | author22=Lindquist, E. | author23=Lucas, S. | author24=Osoegawa, K. | author25=Pennacchio, L. A. | author26=Salamov, A. A. | author27=Satou, Y. | author28=Sauka-Spengler, T. | author29=Schmutz, J. | author30=Shin-i, T. | journal=Nature | volume=453 | issue=7198 | pages=1064–1071 | date=19 June 2008 | bibcode=2008Natur.453.1064P | doi=10.1038/nature06967 | pmid=18563158| doi-access=free }}</ref><ref>{{Cite journal | title=Cyclostome embryology and early evolutionary history of vertebrates | author=Ota, K. G. | author2=Kuratani, S. | journal=Integrative and Comparative Biology | volume=47 | issue=3 | pages=329–337 | date=September 2007 | doi=10.1093/icb/icm022 | pmid=21672842| doi-access=free }}</ref><ref>{{cite journal|vauthors=Delsuc F, Philippe H, Tsagkogeorga G, Simion P, Tilak MK, Turon X, López-Legentil S, Piette J, Lemaire P, Douzery EJ|date=April 2018|title=A phylogenomic framework and timescale for comparative studies of tunicates|journal=BMC Biology|volume=16|issue=1|page=39|doi=10.1186/s12915-018-0499-2|pmc=5899321|pmid=29653534 |doi-access=free }}</ref> Note that [[Placodermi]] is now thought to be [[paraphyletic]].<ref>{{cite journal |last1=Li |first1=Qiang |last2=Zhu |first2=You-an |last3=Lu |first3=Jing |last4=Chen |first4=Yang |last5=Wang |first5=Jianhua |last6=Peng |first6=Lijian |last7=Wei |first7=Guangbiao |last8=Zhu |first8=Min |title=A new Silurian fish close to the common ancestor of modern gnathostomes |journal=Current Biology |date=August 2021 |volume=31 |issue=16 |pages=3613–3620.e2 |doi=10.1016/j.cub.2021.05.053 |pmid=34146483 |s2cid=235477130 |doi-access=free |bibcode=2021CBio...31E3613L }}</ref><ref name="Giles et al 2015">{{closed access}} {{cite journal |last1=Giles |first1=Sam |last2=Friedman |first2=Matt |last3=Brazeau |first3=Martin D. |title=Osteichthyan-like cranial conditions in an Early Devonian stem gnathostome |journal=[[Nature (journal)|Nature]] |volume=520 |issue=7545 |pages=82–85 |date=2015-01-12 |issn=1476-4687 |doi=10.1038/nature14065 |pmid=25581798 |pmc=5536226 |bibcode=2015Natur.520...82G }}</ref> {{clade| style=font-size:85%;line-height:85% |1={{clade |1=[[Cephalochordate|Cephalochordata]] ([[lancelets]]) [[File:Branchiostoma cultellus.jpg|80 px]] |label2=[[Olfactores]] |2={{clade |1=[[Tunicata]] (tunicates: [[sea squirts]], [[Thaliacea|thaliaceans]], [[larvaceans]]) [[File:Salpa scheme.png|60 px]] |label2=[[Vertebrate|Vertebrata]]/ |sublabel2='''Craniata''' |2={{clade |label1=[[Agnatha]]/ |sublabel1=[[Cyclostomes]] |1={{clade | 1=[[Hagfish|Myxini]] ([[hagfishes]]) <span style="{{MirrorH}}">[[File:Cuvier-120-Myxine116.jpg|90 px]]</span> | 5=[[Hyperoartia]]/Petromyzontida ([[lampreys]]) <span style="{{MirrorH}}">[[File:Petromyzon marinus.jpg|80 px]]</span> }} | 2={{clade | 1=†[[Osteostraci]] [[File:Tremataspis NT small.jpg|70 px]] | label2=[[Gnathostomata]] | 2={{clade | state1=double | 1=†[[Placodermi]] [[File:Dunkleosteus terrelli 2023 reconstruction.png|70 px]] | 2={{clade |1=[[Chondrichthyes]] (cartilaginous fishes) <span style="{{MirrorH}}">[[File:White shark (Duane Raver).png|70 px]]</span> |label2=[[Osteichthyes]]/ |sublabel2=[[Euteleostomi]] |2={{clade |1=[[Actinopterygii]] (ray-finned fishes) [[File:Atlantic sturgeon flipped.jpg|70 px]] |2=[[Sarcopterygii]] (lobe-finned fishes and [[tetrapods]]) <span style="{{MirrorH}}">[[File:Coelacanth.png|55 px]]</span> <span style="{{MirrorH}}">[[File:Deinosuchus riograndensis.png|90px]]</span> }} }} }} }} }} }} }} |label1=[[Chordata]]}}
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