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Gynandromorphism
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== As a research tool == Gynandromorphs occasionally afford a powerful tool in genetic, developmental, and behavioral analyses. In ''[[Drosophila melanogaster]]'', for instance, they provided evidence that male courtship behavior originates in the brain,<ref>{{Cite journal|last=Hotta|first=Y, and Benzer, S.|date=1972|title=Mapping of Behaviour in Drosophila mosaics|journal=Nature|volume=240|issue=5383|pages=527β535|doi=10.1038/240527a0|pmid=4568399|bibcode=1972Natur.240..527H|s2cid=4181921}}</ref> that males can distinguish conspecific females from males by the scent or some other characteristic of the posterior, dorsal, [[integument]] of females,<ref>{{Cite journal|last=Nissani|first=M.|date=1975|title=A new behavioral bioassay for an analysis of sexual attraction and pheromones in insects|journal=Journal of Experimental Zoology|volume=192 |issue=2|pages=271β5|doi=10.1002/jez.1401920217|pmid=805823|bibcode=1975JEZ...192..271N }}</ref><ref>{{Cite journal|last=Hotta|first=Y., Benzer, S.|date=1976|title=Courtship in Drosophila mosaics: sex-specific foci for sequential action patterns|journal=Proc Natl Acad Sci U S A|volume=73 |issue=11|pages=4154β4158|doi=10.1073/pnas.73.11.4154|pmid=825859|bibcode=1976PNAS...73.4154H|pmc=431365|doi-access=free}}</ref> that the [[germ cell]]s originate in the posterior-most region of the [[blastoderm]],<ref>{{Cite journal|date=1977|title=Cell lineage analysis of germ cells of Drosophila melanogaster|journal=Nature|volume=265|issue=5596|pages=729β731|doi=10.1038/265729a0|pmid=404558|bibcode=1977Natur.265..729N|last1=Nissani|first1=Moti|s2cid=4146956}}</ref> and that somatic components of the gonads originate in the mesodermal region of the fourth and fifth abdominal segment.<ref>{{Cite journal|last=Szabad|first=Janos, and Nothiger, Rolf|date=1992|title=Gynandromorphs of Drosophila suggest one common primordium for the somatic cells of the female and male gonads in the region of abdominal segments 4 and 5|url=http://dev.biologists.org/content/develop/115/2/527.full.pdf|journal=Development|volume=115|issue=2|pages=527β533|doi=10.1242/dev.115.2.527|pmid=1425338}}</ref>
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