Editing Natural selection (section)

===The modern synthesis===
{{Main|Modern synthesis (20th century)}}<!--of 1918-1932 approx -->

Natural selection relies crucially on the idea of heredity, but developed before the basic concepts of [[genetics]]. Although the [[Moravia]]n monk [[Gregor Mendel]], the father of modern genetics, was a contemporary of Darwin's, his work lay in obscurity, only being rediscovered in 1900.<ref>{{cite web |url=https://www.jic.ac.uk/germplas/PISUM/ZGS4F.HTM |title=Mendel's Peas |last=Ambrose |first=Mike |publisher=Germplasm Resources Unit, [[John Innes Centre]] |location=Norwich, UK |access-date=22 May 2015 |archive-url=https://web.archive.org/web/20160614210558/https://www.jic.ac.uk/germplas/PISUM/ZGS4F.HTM |archive-date=14 June 2016 |url-status=dead }}</ref> With the early 20th-century integration of evolution with [[Mendel's laws]] of inheritance, the so-called [[Modern synthesis (20th century)|modern synthesis]], scientists generally came to accept natural selection.<ref name=Huxley>{{cite book |last=Huxley |first=Julian |author-link=Julian S. Huxley |year=1929–1930 |chapter=The A B C of Genetics |title=The Science of Life |volume=2 |location=London |publisher=[[Amalgamated Press]] |oclc=3171056|title-link=The Science of Life }}</ref><ref>{{cite book |author=National Academy of Sciences |author-link=National Academy of Sciences |year=1999 |title=Science and Creationism: A View from the National Academy of Sciences |url=https://archive.org/details/sciencecreationi0000unse |edition=2nd |location=Washington, DC |publisher=National Academy Press |isbn=978-0-309-06406-4 |oclc=43803228 |url-access=registration }}</ref> The synthesis grew from advances in different fields. Ronald Fisher developed the required mathematical language and wrote ''[[The Genetical Theory of Natural Selection]]'' (1930).<ref name="fisher" /> [[J. B. S. Haldane]] introduced the concept of the "cost" of natural selection.<ref>{{harvnb|Haldane|1932}}</ref><ref>{{cite journal |last=Haldane |first=J. B. S. |author-link=J. B. S. Haldane |date=December 1957 |title=The Cost of Natural Selection |url=http://www.blackwellpublishing.com/ridley/classictexts/haldane2.pdf |journal=[[Journal of Genetics]] |volume=55 |issue=3 |pages=511–524 |doi=10.1007/BF02984069|s2cid=32233460 }}</ref>
[[Sewall Wright]] elucidated the nature of selection and adaptation.<ref>{{cite journal |last=Wright |first=Sewall |author-link=Sewall Wright |year=1932 |title=The roles of mutation, inbreeding, crossbreeding and selection in evolution |url=http://www.blackwellpublishing.com/ridley/classictexts/wright.asp |journal=Proceedings of the VI International Congress of Genetrics |volume=1 |pages=356–366}}</ref>
In his book ''[[Genetics and the Origin of Species]]'' (1937), [[Theodosius Dobzhansky]] established the idea that mutation, [[mutationism|once seen as a rival]] to selection, actually supplied the raw material for natural selection by creating genetic diversity.<ref>{{harvnb|Dobzhansky|1937}}</ref><ref>{{harvnb|Dobzhansky|1951}}</ref>