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===Neoisoptera=== The [[Neoisoptera]], literally meaning "newer termites" (in an evolutionary sense), are a recently coined clade that include families such as the [[Heterotermitidae]], [[Rhinotermitidae]] and [[Termitidae]]. ''Neoisopterans'' have a bifurcated caste development with true workers, and so notably lack pseudergates (except in [[Stylotermitidae]]: see [[#Caste system|below]]). All ''Neoisopterans'' have a fontanelle, which appears as a circular pore or series of pores in a depressed region within the middle of the head. The fontanelle connects to the frontal gland, a novel organ unique to Neoisopteran termites which evolved to excrete an array of defensive chemicals and secretions, and so is typically most developed in the soldier caste.<ref>{{Cite journal |last1=Šobotník |first1=Jan |last2=Bourguignon |first2=Thomas |last3=Hanus |first3=Robert |last4=Sillam-Dussès |first4=David |last5=Pflegerová |first5=Jitka |last6=Weyda |first6=František |last7=Kutalová |first7=Kateřina |last8=Vytisková |first8=Blahoslava |last9=Roisin |first9=Yves |date=2010-12-30 |title=Not Only Soldiers Have Weapons: Evolution of the Frontal Gland in Imagoes of the Termite Families Rhinotermitidae and Serritermitidae |journal=PLOS ONE |volume=5 |issue=12 |pages=e15761 |doi=10.1371/journal.pone.0015761 |pmid=21209882 |pmc=3012694 |bibcode=2010PLoSO...515761S |issn=1932-6203|doi-access=free }}</ref> Cellulose digestion in the family ''Termitidae'' has co-evolved with [[bacteria]]l gut microbiota<ref>{{cite journal | last1 = Kohler | first1 = T | last2 = Dietrich | first2 = C | last3 = Scheffrahn | first3 = RH | last4 = Brune | first4 = A | year = 2012 | title = High-resolution analysis of gut environment and bacterial microbiota reveals functional compartmentation of the gut in wood-feeding higher termites (''Nasutitermes'' spp.) | journal = Applied and Environmental Microbiology | volume = 78 | issue = 13| pages = 4691–4701 | doi=10.1128/aem.00683-12| pmid = 22544239 | pmc = 3370480 | bibcode = 2012ApEnM..78.4691K }}</ref> and many [[Taxon|taxa]] have evolved additional symbiotic relationships such as with the fungus ''[[Termitomyces]]''; in contrast, basal ''Neoisopterans'' and all other ''Euisoptera'' have [[flagellate]]s and [[prokaryote]]s in their hindguts. Extant families and subfamilies are organized as follows:<ref name=":7" /><ref name=":8">{{cite journal | last1 = Hellemans | first1 = Simon | last2 = Rocha | first2 = Mauricio M. | last3 = Wang | first3 = Menglin | last4 = Romero Arias | first4 = Johanna | year = 2024 | title = Genomic data provide insights into the classification of extant termites | journal = Nature Communications | volume = 15 | issue = 1 | page = 6724 | doi=10.1038/s41467-024-51028-y| pmid = 39112457 | pmc = 11306793 }}</ref> : '''Early-Diverging Neoisoptera (Non-Geoisoptera)''' :: Family {{extinct}}[[Archeorhinotermitidae]] <small>Krishna & Grimaldi, 2003</small> :: Family [[Stylotermitidae]] <small>Holmgren & Holmgren, 1917</small> :: Family [[Serritermitidae]] <small>Holmgren, 1910</small> :: Family [[Rhinotermitidae]] <small>Froggatt, 1897</small> :: Family [[Termitogetonidae]] <small>Holmgren, 1910</small> :: Family [[Psammotermitidae]] <small>Holmgren, 1910</small> ::: Subfamily [[Prorhinotermitinae]] <small>Quennedey & Deligne, 1975</small> ::: Subfamily [[Psammotermitinae]] <small>Holmgren, 1910</small> : '''<big>Clade Geoisoptera</big>''' <small>Engel, Hellemans, & Bourguignon, 2024</small> :: Family [[Heterotermitinae|Heterotermitidae]] <small>Froggatt, 1897</small> (<small>=[[Coptotermitinae]] Holmgren, 1910</small>) :: '''Family [[Termitidae]]''' <small>Latreille, 1802</small> ::: Subfamily [[Sphaerotermitinae]] <small>Engel & Krishna, 2004</small> ::: Subfamily [[Macrotermitinae]] <small>Kemner, 1934, nomen protectum [ICZN 2003]</small> ::: Subfamily [[Foraminitermitinae]] <small>Holmgren, 1912</small> ::: Subfamily [[Apicotermitinae]] <small>Grassé & Noirot, 1954 [1955]</small> ::: Subfamily [[Microcerotermitinae]] <small>Holmgren, 1910</small> ::: Subfamily [[Syntermitinae]] <small>Engel & Krishna, 2004</small> ::: Subfamily [[Forficulitermitinae]] <small>Hellemans, Engel, & Bourguignon, 2024</small> ::: Subfamily [[Engelitermitinae]] <small>Romero Arias, Roisin, & Scheffrahn, 2024</small> ::: Subfamily [[Crepititermitinae]] <small>Hellemans, Engel, & Bourguignon, 2024</small> ::: Subfamily [[Protohamitermitinae]] <small>Hellemans, Engel, & Bourguignon, 2024</small> ::: Subfamily [[Cylindrotermitinae]] <small>Hellemans, Engel, & Bourguignon, 2024</small> ::: Subfamily [[Neocapritermitinae]] <small>Hellemans, Engel, & Bourguignon, 2024</small> ::: Subfamily [[Nasutitermitinae]] <small>Hare, 1937</small> ::: Subfamily [[Promirotermitinae]] <small>Hellemans, Engel, & Bourguignon, 2024</small> ::: Subfamily [[Mirocapritermitinae]] <small>Kemner, 1934</small> ::: Subfamily [[Amitermitinae]] <small>Kemner, 1934</small> ::: Subfamily [[Cubitermitinae]] <small>Weidner, 1956</small> ::: Subfamily [[Termitinae]] <small>Latreille, 1802</small>
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