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Apomixis
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====Types of gametophytic apomixis==== Gametophytic apomixis in flowering plants develops in several different ways.<ref name=Nogler>Nogler, G.A. 1984. Gametophytic apomixis. In Embryology of angiosperms. Edited by B.M. Johri. Springer, Berlin, Germany. pp. 475β518.</ref> A megagametophyte develops with an egg cell within it that develops into an embryo through [[parthenogenesis]]. The central cell of the megagametophyte may require fertilization to form the [[endosperm]], '''pseudogamous gametophytic apomixis''', or in '''autonomous gametophytic apomixis''' endosperm fertilization is not required. *In '''diplospory''' (also called '''generative apospory'''), the megagametophyte arises from a cell of the [[archesporium]]. * In '''apospory''' (also called '''somatic apospory'''), the megagametophyte arises from some other (somatic) cell of the nucellus. Considerable confusion has resulted because diplospory is often defined to involve the [[megaspore mother cell]] only, but a number of plant families have a multicellular archesporium and the megagametophyte could originate from another archesporium cell. Diplospory is further subdivided according to how the megagametophyte forms: * ''[[Allium]] odorum''β''A. nutans'' type. The chromosomes double (endomitosis) and then meiosis proceeds in an unusual way, with the chromosome copies pairing up (rather than the original maternal and paternal copies pairing up). * ''[[Taraxacum]]'' type: Meiosis I fails to complete, meiosis II creates two cells, one of which degenerates; three mitotic divisions form the megagametophyte. * ''[[Ixeris]]'' type: Meiosis I fails to complete; three rounds of nuclear division occur without cell-wall formation; wall formation then occurs. * ''[[Blumea]]''β''[[Elymus (plant)|Elymus]]'' types: A mitotic division is followed by degeneration of one cell; three mitotic divisions form the megagametophyte. * ''[[Antennaria]]''β''[[Hieracium]]'' types: three mitotic divisions form the megagametophyte. * ''[[Eragrostis]]''β''[[Panicum]]'' types: Two mitotic division give a 4-nucleate megagametophyte, with cell walls to form either three or four cells.
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