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Formica exsecta
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===Haplodiploidy=== ''F. exsecta'', much like other insects in the order [[Hymenoptera]], have a [[haplodiploid]] sex determination system. An unusual 0.75 [[Coefficient of relationship|relatedness coefficient]] between full haplodiploid sisters is one of the main contributors to the frequency of evolution of eusocial species.<ref>{{cite journal|last=Foster|first=K.|author2=Wenseleers, T.|author3=Ratnieks, F.|title=Kin selection is the key to altruism|journal=Trends in Ecology & Evolution|date=1 February 2006|volume=21|issue=2|pages=57β60|doi=10.1016/j.tree.2005.11.020|pmid=16701471|bibcode=2006TEcoE..21...57F |url=https://lirias.kuleuven.be/handle/123456789/69591|url-access=subscription}}</ref> The queen's eggs that are fertilized grow into [[diploid]] daughters, which contain two pairs of chromosomes, whereas unfertilized eggs produce [[haploid]] males, which only contain the queen's chromosomes. The voluntary fertilization of eggs is done by the egg-laying mother. Therefore, ideally, the queen's best reproductive interest is to lay a larger quantity of eggs or increase the number of eggs that produce individuals that can reproduce themselves. However, female-rich colonies emphasized the production of workers rather than gynes.<ref>{{cite journal|last=Kummerli|first=R.|author2=Keller, L.|title=Reproductive specialization in multiple-queen colonies of the ant ''Formica exsecta''|journal=Behavioral Ecology|date=13 December 2006|volume=18|issue=2|pages=375β383|doi=10.1093/beheco/arl088|url=http://doc.rero.ch/record/302502/files/arl088.pdf|doi-access=free|access-date=18 September 2019|archive-date=16 May 2024|archive-url=https://web.archive.org/web/20240516204816/http://doc.rero.ch/record/302502/files/arl088.pdf|url-status=live}}</ref>
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