Editing Helpers at the nest (section)

=== Reproductive success ===
Effects on reproductive success can be but are not always positive, and the strength of that positive correlation varies by species. Sometimes, the energy benefits to breeders, instead of being invested in improving reproductive performance, are used to offset reductions in reproductive performance.<ref name=":12">{{Cite journal|last1=Manica|first1=Lilian Tonelli|last2=Marini|first2=Miguel Ângelo|date=2012-01-01|title=Helpers at the nest of White-banded Tanager Neothraupis fasciata benefit male breeders but do not increase reproductive success|journal=Journal of Ornithology|language=en|volume=153|issue=1|pages=149–159|doi=10.1007/s10336-011-0718-x|bibcode=2012JOrni.153..149M |s2cid=16198295|issn=2193-7206}}</ref> However, in many species, nestling survival rates are increased from helper investments.<ref name=":2">{{Cite journal|last1=Cusick|first1=Jessica A.|last2=de Villa|first2=Miguel|last3=DuVal|first3=Emily H.|last4=Cox|first4=James A.|date=2018-02-23|title=How do helpers help? Helper contributions throughout the nesting cycle in the cooperatively breeding brown-headed nuthatch|journal=Behavioral Ecology and Sociobiology|language=en|volume=72|issue=3|pages=43|doi=10.1007/s00265-018-2470-1|bibcode=2018BEcoS..72...43C |s2cid=4702911|issn=1432-0762}}</ref> Further reproductive success benefits include better quality offspring,<ref name=":2" /> and nestling [[immune response]].<ref>{{Cite journal|last1=Valencia|first1=Juliana|last2=Solís|first2=Elena|last3=Sorci|first3=Gabriele|last4=de la Cruz|first4=Carlos|date=2006-07-01|title=Positive correlation between helpers at nest and nestling immune response in a cooperative breeding bird|journal=Behavioral Ecology and Sociobiology|language=en|volume=60|issue=3|pages=399–404|doi=10.1007/s00265-006-0179-z|bibcode=2006BEcoS..60..399V |s2cid=1898846|issn=1432-0762|hdl=10630/33525|hdl-access=free}}</ref>