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=== Species === The [[cladogram]] shows the relationships between the tern genera, and the currently recognised species, based on [[mitochondrial DNA]] studies, are listed below:<ref name= bridge/> {{Cladogram|caption=Based on Bridge ''et al'' (2005)<ref name= bridge/> and Černý & Natale (2021)<ref name="Cerny"/> |clades={{clade|style=font-size:80%;line-height:80% |1={{clade |1= ''[[Anous]]'' |2={{clade |1=''[[Gygis]]'' |2={{clade |1=''[[Onychoprion]]'' |2={{clade |1=''[[Sternula]]'' |2={{clade |1=''[[Phaetusa]]'' |2={{clade |1={{clade |1=''[[Gelochelidon]]'' |2=''[[Hydroprogne]]'' }} |2={{clade |1=''[[Larosterna]]'' |2={{clade |1=''[[Chlidonias]]'' |2={{clade |1=''[[Thalasseus]]'' |2=''[[Sterna]]'' }} }} }} }} }} }} }} }} }} }} }} {| class="wikitable collapsible" |- ! Image ! Genus !Species |- |[[File:Anous minutus by Gregg Yan 02.jpg|175px]] |''[[Anous]]'' {{small|Stephens, 1826}} — [[Noddy (tern)|noddies]] | * [[Brown noddy]] (''Anous stolidus'') * [[Black noddy]] (''A. minutus''; sometimes considered to be a subspecies of the lesser noddy)<ref name= harrison388/> * [[Lesser noddy]] (''A. tenuirostris'') * [[Blue noddy]] (''A. cerulea''; formerly placed in a separate genus ''Procelsterna'')<ref name= harrison388/> * [[Grey noddy]] (''A. albivitta''; sometimes considered to be a subspecies of the blue noddy; formerly placed in a separate genus ''Procelsterna'')<ref name= harrison388/> |- |[[File:White tern with fish.jpg|175px]] |''[[Gygis]]'' {{small|Wagler, 1832}} — noddies | * [[White tern]] (''Gygis alba'' also sometimes called white noddy) |- |[[File:Sterna fuscata.JPG|175px]] |''[[Onychoprion]]'' {{small|Wagler, 1832}} — brown-backed terns | * [[Spectacled tern]] (''Onychoprion lunatus'') * [[Bridled tern]] (''O. anaethetus'') * [[Sooty tern]] (''O. fuscatus'') * [[Aleutian tern]] (''O. aleuticus'') |- |[[File:Yellow-billed tern Sternula superciliaris.jpg|175px]] |''[[Sternula]]'' {{small|F. Boie, 1822}} — little white terns | * [[Fairy tern]] (''Sternula nereis'') * [[Damara tern]] (''S. balaenarum'') * [[Little tern]] (''S. albifrons'') * [[Saunders's tern]] (''S. saundersi'' formerly considered to be a subspecies of the little tern)<ref name= harrison382>Harrison (1988) p. 382.</ref> * [[Least tern]] (''S. antillarum'' formerly considered to be a subspecies of the little tern)<ref name= harrison382/> * [[Yellow-billed tern]] (''S. superciliaris'') * [[Peruvian tern]] (''S. lorata'') |- |[[File:Large-billed Tern (Phaetusa simplex), Pantanal, Brazil.jpg|175px]] |''[[Phaetusa]]'' {{small|}} — large-billed terns | * [[Large-billed tern]] (''Phaetusa simplex'') |- |[[File:Sterna-caspia-010.jpg|175px]] |''[[Hydroprogne]]'' {{small|Kaup, 1829}}— Caspian terns | * [[Caspian tern]] (''Hydroprogne caspia'') |- |[[File:Gull-billed Tern in Koonthalulam, India, by Dr. Tejinder Singh Rawal.jpg|175px]] |''[[Gelochelidon]]'' {{small|Brehm, 1830}} — gull-billed terns | * [[Gull-billed tern]] (''Gelochelidon nilotica'') * [[Australian tern]] (''Gelochelidon macrotarsa'') |- |[[File:Larosterna inca3.jpg|175px]] |''[[Larosterna]]'' {{small|}} — Inca terns | * [[Inca tern]] (''Larosterna inca'') |- |[[File:White-winged Tern, Mścichy, Biebrzański Park Narodowy, Polska.jpg|175px]] |''[[marsh tern|Chlidonias]]'' {{small|Rafinesque, 1822}} — marsh terns | * [[Black tern]] (''Chlidonias niger'') * [[White-winged tern]] (or white-winged black tern ''C. leucopterus'') * [[Whiskered tern]] (''C. hybridus'') * [[Black-fronted tern]] (''C. albostriatus'' sometimes placed in ''Sterna'')<ref name = birdlifealbostriata/> |- |[[File:2020-07-18 Thalasseus sandvicensis, St Marys Island, Northumberland 01.jpg|175px]] |''[[Thalasseus]]'' {{small|F. Boie, 1822}} — crested terns | * [[Lesser crested tern]] (''Thalasseus bengalensis'') * [[West African crested tern]] (''Thalasseus albididorsalis'' recently split from ''T. maximus'')<ref name= collinson/> * [[Royal tern]] (''T. maximus'') * [[Greater crested tern]] (''T. bergii'' also known as swift tern) * [[Chinese crested tern]] (''T. bernsteini'') * [[Elegant tern]] (''T. elegans'') * [[Sandwich tern]] (''T. sandvicensis'') * [[Cabot's tern]] (''T. acuflavidus'' recently split from ''T. sandvicensis'')<ref name=ioc>{{cite web| editor1-last=Gill | editor1-first=Frank | editor1-link=Frank Gill (ornithologist) | editor2-last=Donsker | editor2-first=David | year=2019 | title=Sandpipers, snipes, coursers | work=World Bird List Version 9.2 | url=https://www.worldbirdnames.org/bow/sandpipers/ | publisher=International Ornithologists' Union | access-date=25 June 2019 }}</ref> |- |[[File:2014-05-18 Sterna hirundo, Killingworth Lake, Northumberland 02.jpg|175px]] |''[[Sterna]]'' {{small|Linnaeus, 1758}} — large white terns | * [[Forster's tern]] (''Sterna forsteri'') * [[Snowy-crowned tern]] (''S. trudeaui'') * [[Common tern]] (''S. hirundo'') * [[Roseate tern]] (''S. dougallii'') * [[White-fronted tern]] (''S. striata'') * [[Black-naped tern]] (''S. sumatrana'') * [[South American tern]] (''S. hirundinacea'') * [[Antarctic tern]] (''S. vittata'') * [[Kerguelen tern]] (''S. virgata'') * [[Arctic tern]] (''S. paradisaea'') * [[River tern]] (''S. aurantia'') * [[Black-bellied tern]] (''S. acuticauda'' possibly ''Chlidonias'')<ref name= bridge/> * [[White-cheeked tern]] (''S. repressa'' possibly ''Chlidonias'')<ref name= bridge/> |- |} In addition to extant species, the fossil record includes a [[Miocene]] [[chronospecies|palaeospecies]], ''[[Sterna milne-edwardsii]]''.<ref name = fossil>{{cite web | title= Fossil record of the Charadriiformes | url= http://palaeo.gly.bris.ac.uk/Palaeofiles/Fossilgroups/charadriformes/fossils.html | publisher= Palaeobiology and Biodiversity Research Group, University of Bristol | access-date= 13 May 2013 | archive-url= https://web.archive.org/web/20160304023443/http://palaeo.gly.bris.ac.uk/Palaeofiles/Fossilgroups/charadriformes/fossils.html | archive-date= 4 March 2016 | url-status= dead | df= dmy-all }}</ref> The birds in the genus ''[[Anous]]'' are known as noddies, the ''[[Chlidonias]]'' species are the marsh terns,<ref name =HBW/> and all other species comprise the sea terns.<ref name= Barlow>Barlow ''et al.'' (1987) p. 212.</ref><ref name= perrins>Perrins & Arlott (1987) p. 130.</ref>
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