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Amplexus
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===Newts=== In the case of [[newt]]s, the process of amplexus is often observed soon after the newts become seasonally active. In the Western USA, for example, this time is typically soon after the onset of the winter rainy season, when [[intermittent stream]]s and [[vernal pools]] become available as a breeding habitat. The [[rough-skinned newt]] is a specific widespread example of a newt in the western USA that can be observed in quiet stream pools and shallow ponds engaging in amplexus.<ref>C. Michael Hogan (2008) ''[http://www.globaltwitcher.com/artspec_information.asp?thingid=43182 Rough-skinned Newt (Taricha granulosa)] {{webarchive|url=https://web.archive.org/web/20090527153302/http://www.globaltwitcher.com/artspec_information.asp?thingid=43182 |date=2009-05-27 }}'', Globaltwitcher, ed. N. Stromberg</ref> During amplexus in newts, males will typically show the behaviour of tail fanning and chin rubbing which is thought to prompt the mating receptivity of the female newt.<ref name=":11">{{Cite journal|last=Able|first=David J.|date=1999-11-04|title=Scramble competition selects for greater tailfin size in male red-spotted newts (Amphibia: Salamandridae)|journal=Behavioral Ecology and Sociobiology|volume=46|issue=6|pages=423β428|doi=10.1007/s002650050638|bibcode=1999BEcoS..46..423A |s2cid=34638425|issn=0340-5443}}</ref> Studies have shown that male newts who have deeper tail-fins have better control of females during amplexus and are also more successful in catching the females for amplexus.<ref name=":11" /> Additionally, it has been found that the probability of a male newt who has a deeper tail-fin to achieve amplexus is greater than those newts who do not contain a deeper tail-fin, as male newts tend to use their tails during male-male competition.<ref name=":11" /> When a male newt, who is unpaired, encounters a female and male newt engaged in amplexus, the unpaired newt will try to displace the paired male newt by using wrestling tactics.<ref name=":9">{{Cite journal|last=Verrell|first=Paul A.|date=1986|title=Wrestling in the red-spotted newt (Notophthalmus viridescens): resource value and contestant asymmetry determine contest duration and outcome|journal=Animal Behaviour|volume=34|issue=2|pages=398β402|doi=10.1016/s0003-3472(86)80108-7|s2cid=53183312|issn=0003-3472}}</ref> A study examining the wrestling behaviour of newts found that of the observed wrestling encounters, 90% were "won" by the paired male, meaning he would retain the female newt.<ref name=":9" /> The study found that the invading unpaired newt rarely successfully displaces the paired male newt, engaged in amplexus.<ref name=":9" />
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