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Cichlid
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==Distribution and habitat== [[File:Tilapia mariae Australia.jpg|thumb|left|210px|''[[Pelmatolapia mariae]]'', caught on a hook and line, in Australia: Originally from Africa, the species established feral populations in Australia.<ref name=Koehn>{{cite journal |last1=Koehn |first1=John D. |last2=MacKenzie |first2=Rachel F. |title=Priority management actions for alien freshwater fish species in Australia |journal=New Zealand Journal of Marine and Freshwater Research |date=August 2004 |volume=38 |issue=3 |pages=457–472 |doi=10.1080/00288330.2004.9517253 |doi-access=free |bibcode=2004NZJMF..38..457K |s2cid=83569110 }}</ref>]] Cichlids are one of the largest vertebrate families in the world. They are most diverse in Africa and South America. Africa alone is host to at least an estimated 1,600 species.<ref name=Nelson/> [[Central America]] and [[Mexico]] have about 120 species, as far north as the [[Rio Grande]] in [[South Texas]]. [[Madagascar]] has its own distinctive species (''[[Katria katria|Katria]]'', ''[[Oxylapia]]'', ''[[Paratilapia]]'', ''[[Paretroplus]]'', ''[[Ptychochromis]]'', and ''[[Ptychochromoides]]''), only distantly related to those on the African mainland.<ref name="fishbase_CS">{{FishBase family|family=Cichlidae|date=February 2012}}</ref><ref name="Boruchowitz">{{cite book|last=Boruchowitz|first=D. E.|year=2006| title=Guide to Cichlids| publisher=T.F.H. Publications| isbn=978-0-7938-0584-6}}</ref> Native cichlids are largely absent in Asia, except for 9 species in [[Israel]], [[Lebanon]], and [[Syria]] (''[[Astatotilapia flaviijosephi]]'', ''[[Oreochromis aureus]]'', ''[[Oreochromis niloticus|O. niloticus]]'', ''[[Sarotherodon galilaeus]]'', ''[[Coptodon zillii]]'', and ''[[Tristramella]]'' spp.), two in [[Iran]] (''[[Iranocichla]]''), and three in [[India]] and [[Sri Lanka]] (''[[Etroplus]]'' and ''[[Pseudetroplus]]'').<ref name="Nelson"/> If disregarding [[Trinidad and Tobago]] (where the few native cichlids are members of genera that are widespread in the South American mainland), the three species from the genus ''[[Nandopsis]]'' are the only cichlids from the [[Antilles]] in the [[Caribbean]], specifically [[Cuba]] and [[Hispaniola]]. Europe, Australia, Antarctica, and North America north of the Rio Grande drainage have no native cichlids, although in [[Florida]], [[Hawaii]], Japan, northern Australia, and elsewhere, feral populations of cichlids have become established as [[Introduced species|exotics]].<ref name="Koehn"/><ref name="abc.net.au">{{cite web|url=http://www.abc.net.au/farnorth/stories/s1313845.htm |title=Tilapia :: Far North Queensland |author=ABC Far North Queensland |website=[[Australian Broadcasting Corporation]] |access-date=19 April 2007 |archive-url=https://web.archive.org/web/20071017061120/http://abc.net.au/farnorth/stories/s1313845.htm |archive-date=17 October 2007 |url-status=dead}}</ref><ref name="fishbase_convicts">{{cite web|url=http://filaman.uni-kiel.de/Summary/SpeciesSummary.php?id=3615|title=''Archocentrus nigrofasciatus'', Convict cichlid|editor=Froese, R.|editor2=D. Pauly|publisher=FishBase|access-date=29 March 2007|url-status=dead|archive-url=https://web.archive.org/web/20081201110621/http://filaman.uni-kiel.de/Summary/SpeciesSummary.php?id=3615|archive-date=1 December 2008}}</ref><ref name="Yamamoto">{{cite book | author = Yamamoto, M.N.| author2 = Tagawa, A.W.| year = 2000| title = Hawai'i's native and exotic freshwater animals | publisher = Mutual Publishing | location = Honolulu, Hawaii | pages = 200}}</ref><ref name="Page">{{cite book | author = Page, L.M. |author2 = Burr, B.M. | year = 1991| title = A field guide to freshwater fishes of North America north of Mexico| url = https://archive.org/details/fieldguidetofres00lawr | url-access = registration | publisher = Houghton Mifflin Company | location = Boston | pages = [https://archive.org/details/fieldguidetofres00lawr/page/432 432] | isbn = 978-0-395-35307-3}}</ref><ref name="GulfStateMarine">{{cite web|author=University of Southern Mississippi/College of Marine Sciences/Gulf Coast Research Laboratory |title=Fact Sheet for Tilapia zilli (Gervais, 1848) |publisher=Gulf States Marine Fisheries Commission |date=3 August 2005 |url=http://nis.gsmfc.org/nis_factsheet2.php?toc_id=200 |access-date=10 February 2007 |url-status=dead |archive-url=https://web.archive.org/web/20070818063521/http://nis.gsmfc.org/nis_factsheet2.php?toc_id=200 |archive-date=18 August 2007}}</ref><ref name="Fuller">{{cite web | last1=Fuller | first1=Pam L. | first2=Leo G. |last2=Nico | title=Nonindigenous Fishes of Florida – With a Focus on South Florida | publisher=[[United States Geological Survey#Coastal and Marine Science Center|U.S. Department of the Interior, U.S. Geological Survey, Coastal and Marine Science Center]] | date=11 October 2002 | url=http://sofia.usgs.gov/sfrsf/rooms/species/invasive/focus/ |archive-url=https://web.archive.org/web/20030115221654/http://sofia.usgs.gov/sfrsf/rooms/species/invasive/focus/ |archive-date=2003-01-15 |url-status=dead}} </ref> Although no longer present in Europe except as introductions, tilapias are known to have ranged as far north as Italy during the [[Miocene]].<ref name=":02" /> Although most cichlids are found at relatively shallow depths, several exceptions do exist. The deepest known occurrences are ''[[Trematocara]]'' at more than {{convert|300|m|ft|-2|abbr=on}} below the surface in [[Lake Tanganyika]].<ref name=Loiselle1994>Loiselle, Paul (1994). The Cichlid Aquarium, p. 304. Tetra Press, Germany. {{ISBN|978-1564651464}}.</ref> Others found in relatively deep waters include species such as ''[[Alticorpus macrocleithrum]]'' and ''[[Pallidochromis tokolosh]]'' down to {{convert|150|m|ft|-2|abbr=on}} below the surface in Lake Malawi,<ref>{{FishBase| genus = Alticorpus | species= macrocleithrum}}</ref><ref>{{FishBase | genus = Pallidochromis | species= tokolosh }}</ref> and the whitish (non[[Biological pigment|pigmented]]) and blind ''[[Lamprologus lethops]]'', which is believed to live as deep as {{convert|160|m|ft|abbr=on}} below the surface in the [[Congo River]].<ref>Norlander, Britt (20 April 2009). [https://archive.today/20120708055831/http://findarticles.com/p/articles/mi_m1590/is_13_65/ai_n31583235/ Rough waters: one of the world's most turbulent rivers is home to a wide array of fish species. Now, large dams are threatening their future.] ''Science World''</ref> Cichlids are less commonly found in [[Brackish water|brackish]] and [[Seawater|saltwater]] habitats, though many species tolerate brackish water for extended periods; ''[[Mayaheros urophthalmus]]'', for example, is equally at home in freshwater [[marsh]]es and [[mangrove]] swamps, and lives and breeds in saltwater environments such as the mangrove belts around [[barrier island]]s.<ref name="Loiselle"/> Several species of ''[[Tilapia]]'', ''[[Sarotherodon]]'', and ''[[Oreochromis]]'' are [[euryhaline]] and can disperse along brackish coastlines between rivers.<ref name="Nelson"/> Only a few cichlids, however, inhabit primarily brackish or salt water, most notably ''[[Etroplus maculatus]]'', ''[[Etroplus suratensis]]'', and ''[[Sarotherodon melanotheron]]''.<ref name="Schäfer">{{cite book | author = Frank Schäfer | year = 2005 | title = Brackish-Water Fishes | publisher = Aqualog | isbn=978-3-936027-82-2}}{{page needed|date=November 2020}}</ref> The perhaps most extreme habitats for cichlids are the warm [[hypersaline lake]]s where the members of the genera ''[[Alcolapia]]'' and ''[[Danakilia]]'' are found. [[Lake Abaeded]] in [[Eritrea]] encompasses the entire distribution of ''[[Danakilia dinicolai|D. dinicolai]]'', and its temperature ranges from {{convert|29|to|45|C|F}}.<ref>{{cite journal |last1=Stiassny |first1=Melanie L. J. |last2=Marchi |first2=Giuseppe De |last3=Lamboj |first3=Anton |title=A new species of Danakilia (Teleostei, Cichlidae) from Lake Abaeded in the Danakil Depression of Eritrea (East Africa) |journal=Zootaxa |date=29 November 2010 |volume=2690 |issue=1 |pages=43–52 |doi=10.11646/zootaxa.2690.1.4 |s2cid=87705274 |url=https://pdfs.semanticscholar.org/51ff/077f4371672caa31f82b33e56ce59c63368b.pdf |archive-url=https://web.archive.org/web/20200212121742/https://pdfs.semanticscholar.org/51ff/077f4371672caa31f82b33e56ce59c63368b.pdf |url-status=dead |archive-date=12 February 2020 }}</ref> Although the vast majority of Malagasy cichlids are entirely restricted to fresh water, ''[[Ptychochromis grandidieri]]'' and ''[[Paretroplus polyactis]]'' are commonly found in coastal brackish water and apparently are salt tolerant,<ref>{{cite journal |last1=Stiassny |first1=Melanie L.J |last2=Sparks |first2=John S |date=2006 |title=Phylogeny and Taxonomic Revision of the Endemic Malagasy Genus Ptychochromis (Teleostei: Cichlidae), with the Description of Five New Species and a Diagnosis for Katria, New Genus |url=https://www.biodiversitylibrary.org/itempdf/280948 |journal=American Museum Novitates |issue=3535 |page=1 |doi=10.1206/0003-0082(2006)3535[1:PATROT]2.0.CO;2 |s2cid=322493}}</ref><ref>{{cite journal |last1=Sparks |first1=John S. |date=2008 |title=Phylogeny of the Cichlid Subfamily Etroplinae and Taxonomic Revision of the Malagasy Cichlid Genus Paretroplus (Teleostei: Cichlidae) |journal=Bulletin of the American Museum of Natural History |volume=314 |issue=1 |page=1 |doi=10.1206/314.1 |s2cid=84071748}}</ref> as is also the case for ''[[Orange chromide|Etroplus maculatus]]'' and ''[[Green chromide|E. suratensis]]'' from India and Sri Lanka.<ref>{{FishBase|genus=Etroplus|species=maculatus|date=July 2011}}</ref><ref>{{FishBase|genus=Etroplus|species=suratensis|date=July 2011}}</ref>
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