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Triggerfish
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===Spawning and biparental care=== Triggerfish spawning is timed in relation to lunar cycles, tides, and time of changeover of tides. In relation to lunar cycles, eggs are observed 2–6 days before the full moon and 3–5 days before the new moon. In relation to tides, spawning happens 1–5 days before the spring tide. In relation to timing of tides, eggs are observed on days when high tides take place around sunset.<ref name=":7">{{cite journal|last=Gladstone|first=William|date=March 1994|title=Lek-like spawning, parental care and mating periodicity of the triggerfish ''Pseudobalistes flavimarginatus'' (Balistidae).|journal=[[Environmental Biology of Fishes]]|volume=39|issue=3|pages=249–257|doi=10.1007/bf00005127|bibcode=1994EnvBF..39..249G |s2cid=36747250}}</ref> [[File:Titan Triggerfish.jpg|thumb|360px| The [[titan triggerfish]] can move relatively large rocks when feeding and is often followed by smaller fish, in this case [[orange-lined triggerfish]] and [[moorish idol]], that feed on leftovers.]] Male and female triggerfish perform certain prespawning behaviors: blowing and touching.<ref name=":6" /> A male and female blow water on the sandy bottom (usually in the same spot at the same time) and set up their egg site. They touch their abdomens on the bottom as if they are spawning. During actual spawning, eggs are laid on the sandy sea bottom (triggerfish are [[Demersal fish|demersal]] spawners despite their large size). Eggs are scattered and attached to sand particles. Triggerfish eggs are usually very small (diameter of 0.5–0.6 mm)<ref name=":7" /><ref>{{cite journal|last=Kawase|first=Hiroshi|year=2003|title=Maternal egg care in the bridled triggerfish, ''Sufflamen fraenatus'' (Balistidae) at Hachijojima island, Japan.|journal=Natural History Research|volume=7|pages=193–197}}</ref> and are easily spread by waves. After spawning, both the male and female participate in caring for the fertilized eggs (biparental egg care). A female triggerfish stays near the spawning ground, around 5 m off the bottom, and guards the eggs within her territory against intruders. Some common intruders include ''[[Parupeneus multifasciatus]], [[Zanclus cornutus]], [[Prionurus scalprum]]'', and conspecifics. Besides guarding, females roll, fan, and blow water on eggs to provide oxygen to the embryos, thereby inducing hatching.<ref>{{cite web|url=http://www.howfishbehave.ca/pdf/Are%20fishes%20good%20parents.pdf|title=Are fishes good parents?|last=Reebs|first=Stéphan G.|date=2011–2015|publisher=[[Université de Moncton]]|via=howfishbehave.ca|url-status=live|archive-url=https://web.archive.org/web/20160304031133/http://www.howfishbehave.ca/pdf/Are%20fishes%20good%20parents.pdf|archive-date=4 March 2016}}</ref> This behavior of female triggerfish is called "tending", and males rarely perform this behavior. A male triggerfish stays farther above the eggs and guards all the females and eggs in his territory. Males exhibit aggressive behaviors against conspecific males near the boundaries of their territories.
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