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Segnosaurus
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==Paleoenvironment== [[File:Bayan Shireh Formation Fauna.png|thumb|upright=1.5|Dinosaurs from the [[Bayan Shireh Formation]] compared in size (''Segnosaurus'' fifth from right, in dark red)]] Fossils of ''Segnosaurus'' have been recovered from the Bayan Shireh Formation in Mongolia, which has been dated to about 102β86 million years ago during the [[Cenomanian]] to Turonian stages of the Late Cretaceous period, based on [[paleomagnetic]] analysis and [[calcite]] [[UβPb measurements]].<ref name="Magnetic"/><ref name="Formation">{{cite journal |last1=Kurumada |first1=Y. |last2=Aoki |first2=S. |last3=Aoki |first3=K. |last4=Kato |first4=D. |last5=Saneyoshi |first5=M. |last6=Tsogtbaatar |first6=K. |last7=Windley |first7=B. F. |last8=Ishigaki |first8=S. |title=Calcite UβPb age of the Cretaceous vertebrate-bearing Bayn Shire Formation in the Eastern Gobi Desert of Mongolia: usefulness of caliche for age determination |journal=Terra Nova |date=2020 |volume=32 |issue=4 |pages=246β252 |doi=10.1111/ter.12456|bibcode=2020TeNov..32..246K |doi-access=free }}</ref> The remains were found in poorly cemented, gray sands containing intraformational [[Conglomerate (geology)|conglomerates]], gravel, and gray claystone.<ref name=Dinosauria1/> The Bayan Shireh Formation overlies the [[Baruunbayan Formation]] and underlies the Javkhlant Formation. The sediments of these formations were deposited by [[meandering rivers]] and lakes on an [[alluvial plain]] (flat land consisting of sediments deposited by highland rivers) with a [[semi-arid]] climate.<ref name="Arid">{{cite journal |last1=Khand |first1=Y. |last2=Badamgarav |first2=D. |last3=Ariunchimeg |first3=Y. |last4=Barsbold |first4=R. |title=Cretaceous system in Mongolia and its depositional environments |journal=Developments in Palaeontology and Stratigraphy |date=2000 |volume=17 |pages=49β79 |doi=10.1016/S0920-5446(00)80024-2 |publisher=Elsevier|isbn=9780444502766 }}</ref><ref name="Trackway"/> Therizinosaurs were the most abundant theropods in the Bayan Shireh Formation in terms of [[biodiversity]]; in addition to ''Segnosaurus'', members of the group included ''Erlikosaurus'', ''Enigmosaurus'', and possibly a fourth type.<ref name="Trackway">{{cite journal |last1=Lee |first1=Yu.-N. |last2=Lee |first2=H.-J. |last3=Kobayashi |first3=Y. |last4=Paulina-Carabajal |first4=A. |last5=Barsbold |first5=R. |last6=Fiorillo |first6=A. R. |last7=Tsogtbaatar |first7=K. |title=Unusual locomotion behaviour preserved within a crocodyliform trackway from the Upper Cretaceous Bayanshiree Formation of Mongolia and its palaeobiological implications |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |date=2019 |volume=533 |pages=109239 |doi=10.1016/j.palaeo.2019.109239|bibcode=2019PPP...53309239L |s2cid=197584839 }}</ref><ref name="Didactyl">{{cite conference |last1=Kobayashi |first1=Y. |last2=Tsogtbaatar |first2=C. |last3=Tsogtbaatar |first3=K. |last4=Barsbold |first4=R. |title=A new therizinosaur with functionally didactyl hands from the Bayanshiree Formation (Cenomanian-Turonian), Omnogovi Province, southeastern Mongolia |journal=Journal of Vertebrate Paleontology |date=2015 |page=157 |location=Dallas}}</ref> Other theropods included the tyrannosaur ''[[Alectrosaurus]]'', the ornithomimid ''[[Garudimimus]]'', and the dromaeosaur ''[[Achillobator]]''.<ref name="Magnetic"/><ref name=Perle1999>{{cite journal|last1=Perle|first1=A.|last2=Norell|first2=M. A.|last3=Clark|first3=J.|title=A new maniraptoran theropod β ''Achillobator giganticus'' (Dromaeosauridae) β from the Upper Cretaceous of Burkhant, Mongolia|journal=Contributions from the Geology and Mineralogy Chair, National Museum of Mongolia|date=1999|issue=101|pages=1β105|oclc=69865262}}</ref> Other dinosaurs included the ankylosaur ''[[Talarurus]]'',<ref name="Magnetic"/> the hadrosaur ''[[Gobihadros]]'',<ref name="Gobihadros">{{cite journal |last1=Tsogtbaatar |first1=K. |last2=Weishampel |first2=D. B. |last3=Evans |first3=D. C. |last4=Watabe |first4=M. |last5=Mpodozis |first5=J. M. |title=A new hadrosauroid (Dinosauria: Ornithopoda) from the Late Cretaceous Baynshire Formation of the Gobi Desert (Mongolia) |journal=PLOS ONE |date=2019 |volume=14 |issue=4 |pages=e0208480 |doi=10.1371/journal.pone.0208480|pmid=30995236 |pmc=6469754 |bibcode=2019PLoSO..1408480T |doi-access=free }}</ref> the sauropod ''[[Erketu]]'',<ref>{{cite journal |last1=Ksepka |first1=D. T. |last2=Norell |first2=M. |title=''Erketu ellisoni'', a long-necked sauropod from Bor GuvΓ© (Dornogov Aimag, Mongolia)|journal=American Museum Novitates|issue=3508 |pages=1β16|date=2006 |hdl=2246/5783 |doi=10.1206/0003-0082(2006)3508[1:EEALSF]2.0.CO;2 |s2cid=86032547 |url=https://www.biodiversitylibrary.org/bibliography/169093 }}</ref> and the ceratopsian ''[[Graciliceratops]]''.<ref>{{cite book |last1=Sereno |first1=P. C. |chapter=The fossil record, systematics and evolution of pachycephalosaurs and ceratopsians from Asia |year=2000 |title=The Age of Dinosaurs in Russia and Mongolia|pages=480β516 |isbn=978-0-521-54582-2 |location=Cambridge|publisher=Cambridge University Press| editor1 =Benton, M. J.| editor2 =Shishkin, M. A.| editor3 =Unwin, D. M.| editor4 =Kurochkin, E. N.}}</ref> Dinosaur eggs, some of which were identified as Dendroolithidae, as well as footprints of dinosaurs and [[crocodyliforms]], have also been found. The formation is distinctive for its variety and abundance of turtles, and invertebrates include [[ostracods]] and freshwater molluscs.<ref name="Magnetic">{{cite journal |last1=Hicks |first1=J. F. |last2=Brinkman |first2=D. L. |last3=Nichols |first3=D. J. |last4=Watabe |first4=M. |title=Paleomagnetic and palynologic analyses of Albian to Santonian strata at Bayn Shireh, Burkhant, and Khuren Dukh, eastern Gobi Desert, Mongolia |journal=Cretaceous Research |date=1999 |volume=20 |issue=6 |pages=829β850 |doi=10.1006/cres.1999.0188|bibcode=1999CrRes..20..829H }}</ref><ref name=Dinosauria1/><ref name="Trackway"/> The Bayan Shireh Formation is possibly coeval with the [[Iren Dabasu Formation]] of the [[Inner Mongolia]] region of China, from where therizinosaur fossils similar to those of ''Segnosaurus'' and ''Erlikosaurus'' have also been found.<ref name="IrenDabasu">{{cite journal |last1=Currie |first1=P. J. |last2=Eberth |first2=D. A. |title=Palaeontology, sedimentology and palaeoecology of the Iren Dabasu Formation (Upper Cretaceous), Inner Mongolia, People's Republic of China |journal=Cretaceous Research |date=1993 |volume=14 |issue=2 |pages=127β144 |doi=10.1006/cres.1993.1011|bibcode=1993CrRes..14..127C }}</ref>
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