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Diplodocidae
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==Classification== [[File:Dinosaurs in Their Time- Jurassic Era Room (2705542612).jpg|left|thumb|Holotype specimens of ''Diplodocus carnegii'' and ''Apatosaurus louisae'' at [[Carnegie Museum of Natural History]]]] Diplodocidae was the third name given to what is now recognized as the single family of long-necked, whip-tailed sauropods. [[Edward Drinker Cope]] named the family '''Amphicoeliidae''' in 1878 for his genus ''[[Amphicoelias]]'', sometimes considered a diplodocid.<ref name="Carpenter2006"/> However, the name Amphicoeliidae did not come into wider use and was not used in the scientific literature after 1899, making it a ''[[nomen oblitum]]'' ("forgotten name") according to the [[International Code of Zoological Nomenclature|ICZN]], preventing it from displacing the name Diplodocidae as a senior synonym.<ref name="Olshevsky1991"/> More recent studies have also shown that ''Amphicoelias'' itself does not belong to this family, but is instead a more primitive diplodocoid.<ref name="Whitlock2011"/> A similar situation occurred for the family name '''Atlantosauridae''', named by [[Othniel Charles Marsh]] in 1877, and which Hay argued had priority over Amphicoelidae.<ref name="Hay1902"/> George Olshevsky declared Atlantosauridae a ''nomen oblitum'' in 1991, though scientists such as Steel and Nowinski had treated Atlantosauridae as a valid name as late as 1971, and the former even added a subfamily, '''Atlantosaurinae'''.<ref name="Olshevsky1991"/><ref name="Nowinski1971"/> Some dinosaurs have been considered diplodocids in the past but have not been found to be members of that group in later, larger analyses of the family's relationships. ''[[Australodocus]]'', for example, was initially described as a diplodocid, but may actually have been a [[Macronaria]]n.<ref name="Whitlock2011"/><ref name="Remes2007"/> ''Amphicoelias'' was traditionally considered a diplodocid due to its similar anatomy, but phylogenetic studies showed it to be a more basal member of the Diplodocoidea.<ref name="Whitlock2011"/> The relationships of species within Diplodocidae has also been subject to frequent revision. A study by Lovelace, Hartman and Wahl in 2008 found that ''[[Suuwassea]]'' and ''Supersaurus'' were relatives of ''Apatosaurus'', within the subfamily Apatosaurinae. However, a subsequent analysis by Whitlock in 2011 showed that ''Supersaurus'' is slightly closer to ''Diplodocus'' than to ''Apatosaurus'', and that ''Suuwassea'' is actually a primitive [[Dicraeosauridae|dicraeosaurid]].<ref name="Whitlock2011"/> [[File:Apatosaurus Clean.png|thumb|Cast skeleton of ''Brontosaurus parvus'' based on the University of Wyoming specimen produced by [[Triebold Paleontology Incorporated]]]] The subfamily '''Diplodocinae''', was erected to include ''Diplodocus'' and its closest relatives, including ''[[Barosaurus]]''.<ref name="TaylorNaish2005"/><ref name="Harris2006"/> The Portuguese ''[[Dinheirosaurus]]'' and the African ''[[Tornieria]]'' have also been identified as close relatives of ''Diplodocus'' by some authors.<ref name="BonaparteMateus1999"/><ref name="Rauhut2005"/> [[Cladogram]] of the Diplodocidae after Tschopp, Mateus, and Benson (2015).<ref name="Tschopp2015"/> {{clade| style=font-size:85%;line-height:85% |label1='''Diplodocidae''' |1={{clade |1=''[[Amphicoelias altus]]'' |2={{clade |label1=[[Apatosaurinae]] |1={{clade |1=Unnamed species |2={{clade |1={{clade |1=''[[Apatosaurus ajax]]'' |2=''[[Apatosaurus louisae]]'' }} |2={{clade |1=''[[Brontosaurus excelsus]]'' |2={{clade |1=''[[Brontosaurus yahnahpin]]'' |2=''[[Brontosaurus parvus]]'' }} }} }} }} |label2=[[Diplodocinae]] |2={{clade |1=Unnamed species |2={{clade |1=''[[Tornieria africana]]'' |2={{clade |1={{clade |1=''[[Supersaurus lourinhanensis]]'' |2=''[[Supersaurus vivianae]]'' }} |2={{clade |1=''[[Leinkupal laticauda]]'' |2={{clade |1=''[[Galeamopus hayi]]'' |2={{clade |1={{clade |1=''[[Diplodocus carnegii]]'' |2=''[[Diplodocus hallorum]]'' }} |2={{clade |1=''[[Kaatedocus siberi]]'' |2=''[[Barosaurus lentus]]'' }} }} }} }} }} }} }} }} }} }}
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