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Bean weevil
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{{short description|Subfamily of beetles}} {{Automatic taxobox | name = Bean weevils | oldest_fossil = Cenomanian | image = Bruchinae in Xinjiang.jpg | image_caption = | taxon = Bruchinae | authority = [[Pierre André Latreille|Latreille]], 1802<ref>{{ITIS |id=678800 |taxon=Bruchinae}}</ref> | diversity_link = #Genera | diversity = About 1,650 species in 70 genera | diversity_ref = <ref>{{cite book|first=Geoffrey|last=Morse|year=2014|chapter=2.7.1 Bruchinae Latreille, 1802|pages=189–198|editor-last1=Leschen|editor-first1=R.A.B.|editor-last2=Beutel|editor-first2=R.G.|title=Handbook of Zoology. Arthropoda: Insecta: Coleoptera: Volume 3: Morphology and Systematics (Phytophaga)|publisher=Walter de Gruyter|place=Berlin - Boston|isbn=978-3-11-027370-0|doi=10.1515/9783110274462.189}}</ref> | subdivision_ranks = Tribes and subtribes | subdivision_ref = <ref>{{cite journal|first1=Patrice|last1=Bouchard|first2=Yves|last2=Bousquet|first3=Anthony E.|last3=Davies|first4=Miguel A.|last4=Alonso-Zarazaga|first5=John F.|last5=Lawrence|first6=Chris H. C.|last6=Lyal|first7=Alfred F.|last7=Newton|first8=Chris A. M.|last8=Reid|first9=Michael|last9=Schmitt|first10=S. Adam|last10=Ślipiński|first11=Andrew B. T.|last11=Smith|year=2011|title=Family-group names in Coleoptera (Insecta)|journal=[[ZooKeys]]|issue=88|pages=1–972|doi=10.3897/zookeys.88.807|doi-access=free|pmc=3088472|pmid=21594053}}</ref> | subdivision = * [[Amblycerini]] ** [[Amblycerina]] ** [[Spermophagina]] * [[Bruchini]] ** [[Acanthoscelidina]] (= [[Bruchidiina]]) ** [[Bruchina]] ** [[Megacerina]] * [[Eubaptini]] * [[Kytorhinini]] * [[Pachymerini]] ** [[Caryedontina]] ** [[Caryopemina]] ** [[Pachymerina]] * [[Rhaebini]] * †[[Myanmaropini]]<ref name="Legalovetal2020"/> | synonyms = Lariidae <small>Bedel, 1901</small> }} [[Image:Bonenkever Acanthoscelides obtectus.jpg|thumb|right|Damage to beans by larvae of the common bean weevil, ''[[Acanthoscelides obtectus]]'']] The '''bean weevils''' or '''seed beetles''' are a [[subfamily]] ('''Bruchinae''') of [[weevil]]s, which in turn are a subset of [[beetle]]s. The Bruchinae now placed in the family [[Chrysomelidae]], though they have historically been treated as a separate family. The subfamily includes about 1,650 species and are found worldwide. Bean weevils are generally compact and oval in shape, with small heads somewhat bent under. Sizes range from {{Convert|1|-|22|mm|abbr=out}} for some tropical species. Colors are usually black or brown, often with mottled patterns. Although their [[Mandible (insect mouthpart)|mandible]]s may be elongated, they do not have the long snouts characteristic of other weevils. A defining characteristic of this subfamily is that the [[Elytron|elytra]] don't quite reach the tip of their [[Abdomen (insect anatomy)|abdomen]]s (as seen in the header image). The Bruchinae are [[granivore]]s, and typically infest various kinds of [[seed]]s or [[bean]]s (hence their [[common name]]s), living most of their lives inside a single seed;{{Citation needed|date=May 2025}} Host plants tend to be [[legume]]s, but species will also be found in [[Convolvulaceae]], [[Arecaceae]], and [[Malvaceae]]. The adults deposit eggs on seeds of these plants, then the larvae chew their way into the seed. When ready to [[pupa]]te, the larvae typically cut an exit hole, then return to their feeding chamber. Due to this, several species are considered [[Pest (organism)|pest]]s. The adult weevils have a habit of feigning death and dropping from a plant when disturbed. Several species are native to Great Britain, but there are also records of several [[introduced species]] from stored products in [[warehouse]]s and [[dwelling]]s, although these species cannot proliferate outside of [[Heating, ventilation, and air conditioning|heated buildings]] in that climate.{{Relevance inline|date=May 2025|reason=Why does the UK have to be discussed and not other countries?}} ==Genera== This list of genera uses the new [[Taxonomy|classification]] scheme, treating the bean weevils as a subfamily with six extant tribes, eight or nine subtribes, and one extinct tribe. The former names and ranks for the tribes and subtribes under the old classification, treating the been weevils as a family, are given in parentheses (except for Myanmaropini, which was established only for the new classification).<ref>{{cite web|title=Species Information|url=http://home.sandiego.edu/~gmorse/Species.htm|website=BRUCHBASE|access-date=12 August 2021}}</ref><ref>{{cite web|title=Coleoptera: Bruchidae|url=http://www.coleoptera.org/p1541.htm|website=Coleoptera.org|access-date=29 July 2021|archive-date=15 September 2009|archive-url=https://web.archive.org/web/20090915003851/http://coleoptera.org/p1541.htm|url-status=dead}}</ref> * Tribe [[Amblycerini]] Bridwell, 1932 (= Subfamily Amblycerinae) ** Subtribe [[Amblycerina]] Bridwell, 1932 (= Tribe Amblycerini) *** ''[[Amblycerus]]'' Thunberg, 1815 ** Subtribe [[Spermophagina]] Borowiec, 1987 (= Tribe Spermophagini) *** ''[[Spermophagus]]'' Schoenherr, 1833 *** ''[[Zabrotes]]'' Horn, 1885 * Tribe [[Bruchini]] Latreille, 1802 (= Subfamily Bruchinae) ** Subtribe [[Acanthoscelidina]] Bridwell, 1946 (= Tribe Acanthoscelidini) *** ''[[Abutiloneus]]'' Bridwell, 1946 *** ''[[Acanthoscelides]]'' Schilsky, 1905 *** ''[[Algarobius]]'' Bridwell, 1946 *** ''[[Althaeus]]'' Bridwell, 1946 *** ''[[Bonaerius]]'' Bridwell, 1952 *** ''[[Caryedes]]'' Hummel, 1827 *** ''[[Cosmobruchus]]'' Bridwell, 1931 *** ''[[Ctenocolum]]'' Kingsolver & Whitehead, 1974 *** ''[[Dahlibruchus]]'' Bridwell, 1931 *** ''[[Gibbobruchus]]'' Pic, 1913 *** ''[[Lithraeus]]'' Bridwell, 1952 *** ''[[Margaritabruchus]]'' Romero & Johnson, 2001 *** ''[[Meibomeus]]'' Bridwell, 1946 *** ''[[Megasennius]]'' Whitehead & Kingsolver, 1975 *** ''[[Merobruchus]]'' Bridwell, 1946 *** ''[[Mimosestes]]'' Bridwell, 1946 *** ''[[Neltumius]]'' Bridwell, 1946 *** ''[[Neobruchidius]]'' Johnson & Romero, 2006 *** ''[[Palpibruchus]]'' Borowiec, 1987 *** ''[[Pectinibruchus]]'' Kingsolver, 1967 *** ''[[Penthobruchus]]'' Kingsolver, 1973 *** ''[[Pseudopachymerina]]'' Zacher, 1952 *** ''[[Pygiopachymerus]]'' Pic, 1911 *** ''[[Rhipibruchus]]'' Bridwell, 1932 *** ''[[Scutobruchus]]'' Kingsolver, 1968 *** ''[[Sennius]]'' Bridwell, 1946 *** ''[[Spatulobruchus]]'' Borowiec, 1987 *** ''[[Stator (beetle)|Stator]]'' Bridwell, 1946 *** ''[[Stylantheus]]'' Bridwell, 1946 ** Subtribe [[Bruchidiina]] Bridwell, 1946 (= Tribe Bruchidiini){{NoteTag|This subtribe is sometimes combined with Acanthoscelidina.}} *** ''[[Acanthobruchidius]]'' Borowiec, 1980 *** ''[[Borowiecius]]'' Anton, 1994 *** ''[[Bruchidius]]'' Schilsky, 1905 *** ''[[Callosobruchus]]'' Pic, 1902 *** ''[[Conicobruchus]]'' Decelle, 1951 *** ''[[Decellebruchus]]'' Borowiec, 1987 *** ''[[Horridobruchus]]'' Borowiec, 1984 *** ''[[Kingsolverius]]'' Borowiec, 1987 *** ''[[Megabruchidius]]'' Borowiec, 1984 *** ''[[Palaeoacanthoscelides]]'' Borowiec, 1985 *** ''[[Parasulcobruchus]]'' Anton, 1999 *** ''[[Pygobruchidius]]'' Pic, 1951 *** ''[[Salviabruchus]]'' Decelle, 1982 *** ''[[Specularius]]'' Bridwell, 1938 *** ''[[Sulcobruchus]]'' Chûjô, 1937 *** ''[[Tuberculobruchus]]'' Decelle, 1951 ** Subtribe [[Bruchina]] Latreille, 1802 (= Tribe Bruchini) *** ''[[Bruchus]]'' Linnaeus, 1767 ** Subtribe [[Megacerina]] Bridwell, 1946 (= Tribe Megacerini) *** ''[[Megacerus]]'' Fåhraeus, 1839 * Tribe [[Eubaptini]] Bridwell, 1932 (= Subfamily Eubaptinae) ** ''[[Eubaptus]]'' Lacordaire, 1845 * Tribe [[Kytorhinini]] Bridwell, 1832 (= Subfamily Kytorhininae) ** ''[[Kytorhinus]]'' Fischer von Waldheim, 1809 * Tribe [[Pachymerini]] Bridwell, 1929 (= Subfamily Pachymerinae) ** Subtribe [[Caryedontina]] Bridwell, 1929 (= Tribe Caryedontini) *** ''[[Aforedon]]'' Decelle, 1965 *** ''[[Caryedon]]'' Schoenherr, 1823 *** ''[[Caryotrypes]]'' Decelle, 1968 *** ''[[Exoctenophorus]]'' Decelle, 1968 *** ''[[Mimocaryedon]]'' Decelle, 1968 ** Subtribe [[Caryopemina]] Bridwell, 1929 (= Tribe Caryomepini) *** ''[[Caryopemon]]'' Jekel, 1855 *** ''[[Diegobruchus]]'' Pic, 1913 *** ''[[Protocaryopemon]]'' Borowiec, 1987 ** Subtribe [[Pachymerina]] Bridwell, 1929 (= Tribe Pachymerini) *** ''[[Butiobruchus]]'' Prevett, 1966 *** ''[[Caryobruchus]]'' Bridwell, 1929 *** ''[[Caryoborus]]'' Schoenherr, 1833 *** ''[[Pachymerus]]'' Thunberg, 1805 ***†''[[Mesopachymerus]]'' Poinar, 2005 [[Canadian amber]], [[Campanian]] * Tribe [[Rhaebini]] Chapuis, 1874 (= Subfamily Rhaebinae) ** ''[[Rhaebus (beetle)|Rhaebus]]'' Fischer von Waldheim, 1824 * Tribe †[[Myanmaropini]] Legalov ''et al.'', 2020 **†''[[Myanmarops]]'' Legalov ''et al.'', 2020<ref name="Legalovetal2020">{{Cite journal|last1=Legalov|first1=Andrei A.|last2=Kirejtshuk|first2=Alexander G.|last3=Anokhin|first3=Boris A.|date=March 2020|title=The oldest seed beetle (Coleoptera; Chrysomelidae: Bruchinae) from Upper Cretaceous amber of northern Myanmar with description of new tribe, genus and species|url=https://linkinghub.elsevier.com/retrieve/pii/S0195667119302794|journal=Cretaceous Research|language=en|volume=107|pages=104283|doi=10.1016/j.cretres.2019.104283|s2cid=210632430 |url-access=subscription}}</ref> [[Burmese amber]], Myanmar, [[Cenomanian]] ==Notes== {{reflist|group=note}} ==References== {{Reflist}} * [[John M. Kingsolver]], "Bruchidae", in [[Ross H. Arnett, Jr.]] and [[Michael C. Thomas]], ''[[American Beetles]]'' (CRC Press, 2002), vol. 2 (note that this reference itself indicates the group is a subfamily, and the change was made only after the chapter was written) ==External links== {{Wikispecies|Bruchinae}} {{Commons category|Bruchinae}} * [http://home.sandiego.edu/~gmorse/BRUCHBASE.htm BRUCHBASE] {{Taxonbar|from1=Q217617|from2=Q13167641}} [[Category:Bruchinae| ]] [[Category:Storage pests]]
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