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{{Short description|Genus of plants in Combretaecae family}} {{Hatnote|"Schousboea" redirects here; this is also synonym of ''[[Alchornea]]''.}} {{Use dmy dates|date=August 2024}} {{Automatic taxobox |image = Combretum collinum Bild0860.jpg |image_caption = ''[[Combretum collinum]]'' with fruit |taxon = Combretum |authority = [[Pehr Löfling|Loefl.]] |subdivision_ranks = Species |subdivision = About 3272 see [[#Selected species|text]] |synonyms = *''Aetia'' {{small|Adans.}} *''Bucholzia'' {{small|Stadtm. ex Willemet}} *''Bureava'' {{small|Baill.}} *''Cacoucia'' {{small|Aubl.}} *''Calopyxis'' {{small|Tul.}} *''Campylochiton'' {{small|Welw. ex Hiern}} *''Campylogyne'' {{small|Welw. ex Hemsl.}} *''Chrysostachys'' {{small|Pohl}} *''Codonocroton'' {{small|E.Mey. ex Engl. & Diels}} *''Cristaria'' {{small|Sonn.}} *''Embryogonia'' {{small|Blume}} *''Forsgardia'' {{small|Vell.}} *''Gonocarpus'' {{small|Ham.}} *''Grislea'' {{small|L.}} *''Hambergera'' {{small|Scop.}} *''Kleinia'' {{small|Crantz}} *''Meiostemon'' {{small|Exell & Stace}} *''Mekistus'' {{small|Lour. ex Gomes Mach.}} *''Physopodium'' {{small|Desv.}} *''Poivrea'' {{small|Comm. ex DC.}} *''Quisqualis'' {{small|L.}} *''Schousboea'' {{small|Willd.}} *''Seguiera'' {{small|Rchb. ex Oliv.}} *''Sheadendron'' {{small|G.Bertol.}} *''Sphalanthus'' {{small|Jack}} *''Thiloa'' {{small|Eichler}} *''Udani'' {{small|Adans.}} |synonyms_ref =<ref name="powo">{{cite web |title=Combretum Loefl. |url=http://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:331508-2 |website=Plants of the World Online |publisher=Royal Botanic Gardens, Kew |access-date=11 April 2021}}</ref> }} '''''Combretum''''', the '''bushwillows''' or '''combretums''', make up the [[type genus]] of the family [[Combretaceae]]. The [[genus]] comprises about 272 [[species]] of trees and shrubs, most of which are native to [[Tropics|tropical]] and southern Africa, about 5 to Madagascar, but there are others that are native to tropical Asia, [[New Guinea]] and the [[Bismarck Archipelago]], Australia, and [[Neotropics|tropical America]]. Around 17 species in the genus '''Quisqualis''' are very similar to Combretum and are now classified as species of the genus.<ref>Jordaan, M., Van Wyk, A. E., & Maurin, O. (2011). Generic status of Quisqualis (Combretaceae), with notes on the taxonomy and distribution of Q. parviflora. Bothalia, 41(1), 161-169.</ref> Though somewhat reminiscent of [[willow]]s (''Salix'') in their [[Morphology (biology)|habitus]], they are not particularly close relatives of these. ==Ecology== [[Image:Combretum albidum (Piluki) in Kinnarsani WS, AP W2 IMG 5783.jpg|left|thumb|196px|''[[Combretum albidum]]'' in [[Kinnerasani Wildlife Sanctuary]], [[Andhra Pradesh]], [[India]] ]] Bushwillow trees often are important plants in their [[habitat]]. [[Savannah]]s in Africa, in particular those growing on [[granite|granitic]] soils, are often dominated by ''Combretum'' and its close relative ''[[Terminalia (plant)|Terminalia]]''. For example, ''C. apiculatum'' is a notable tree in the [[Angolan mopane woodlands]] ecoregion in the [[Kunene River]] basin in southern Africa.<ref>C.Michael Hogan. 2012. [http://www.eoearth.org/article/Kunene_River?topic=78166 ''Kunene River''. Topic ed. P.Saundry. Encyclopedia of Earth. National Council for Science and the Environment. Washington DC.] </ref> Other species of this genus are a major component of [[Southwestern Amazonian moist forests]]. This genus contains several species that are [[pollinated]] by [[mammal]]s other than [[bat]]s, which is quite rare indeed.<ref name="Carthew1997">{{cite journal|last=Carthew|first=S|year=1997|title=Non-flying mammals as pollinators|journal=Trends in Ecology & Evolution|volume=12|issue=3|pages=104–108|issn=0169-5347|doi=10.1016/S0169-5347(96)10067-7|pmid=21237993|bibcode=1997TEcoE..12..104C}}</ref> But most species are more conventionally pollinated by [[insect]]s or [[bird]]s.{{citation needed|date=July 2012}} ''[[Typhlodromus combretum]]'', a [[mite]] of the family [[Phytoseiidae]], was discovered on a bushwillow plant and is named after this genus. Other [[herbivore]]s that eat ''Combretum'' foliage include the [[caterpillar]]s of the [[Brown Awl]] (''Badamia exclamationis'') which is found on ''[[Combretum albidum|C. albidum]], [[Combretum latifolium|C. latifolium]]'' and ''[[Combretum ovalifolium|C. ovalifolium]]''; those of the [[Orange-tailed Awl]] (''Bibasis sena'') are recorded from ''[[Combretum extensum|C. extensum]]'' and ''C. latifolium''.{{citation needed|date=July 2012}} ==Use by humans== [[File:Combretum aculeatum MS 10043.jpg|thumb|right|''[[Combretum aculeatum]]'' [[inflorescence]]]] [[File:Combretum paniculatum MS 10060.jpg|thumb|right|''[[Combretum paniculatum]]'']] [[File:Combretum acutum MHNT.BOT.2009.16.15.jpg|thumb|''Combretum acutum'' – [[MHNT]]]] Several species are used in [[African traditional medicine|African]] or [[Ayurveda|Indian]] [[herbal medicine]].{{additional citation needed|date=August 2023}} Several species of this genus in [[Madagascar]] known collectively as ''voantamenaka'' or ''voatamenaka'' (from ''voa'' 'fruit' and ''menaka'' 'oil') are used in traditional Malagasy medicine as [[deworming]] remedies.<ref name="Boiteau">{{Cite dictionary |title=''Calopyxis'' |dictionary=Dictionnaire des noms malgaches de végétaux |publisher=Editions Alzieu |date=1999 |author-link=Pierre Boiteau |volume=I |page=196 |language=fr |first1=Pierre |last1=Boiteau |url=https://malagasyword.org/bins/teny2/voantamenaka}}</ref> The class of chemical compounds known as [[combretastatin]]s were first isolated from [[Combretum afrum|South African bushwillow]] (''Combretum afrum''), from which they get their name. One synthetic [[derivative (chemistry)|derivative]], [[fosbretabulin disodium]] (combretastatin A4 phosphate), underwent preliminary study for the treatment of [[anaplastic thyroid cancer]],<ref>{{cite web |url=http://clinicaltrials.gov/ct2/show/NCT00060242 | title = Combretastatin A4 However there is the possibility that a modified version of an isomer of aombretastatin-A4 can be rendered effective. After replacing adjacent carbon atoms with nitrogen atoms, using blue light, the isomer can be switched to a form that attacks microtubules of cancer cells. So far this has been tested in vitro with breast cancer cells. Phosphate in Treating Patients With Advanced Anaplastic Thyroid Cancer | date = 10 June 2010 | publisher = clinicaltrials.gov | last1 = Savvides | first1 = Panayiotis }}</ref> but it was not effective enough to progress to more advanced trials. ''[[Combretum molle|C. molle]]'' is also recorded to contain [[antioxidant]]s such as [[punicalagin]],{{Citation needed|date=February 2012}} which is also found in [[pomegranate]]s (''Punica granatum''), a somewhat related plant. The [[botanist]] [[George Don]] studied this genus extensively. The [[Luvuvhu River]] in southern Africa was named after the river's bushwillows (''[[Combretum erythrophyllum|C. erythrophyllum]]''), locally known as ''muvuvhu''. ==Species== As of April 2021, there are 272 accepted species of ''Combretum'':<ref name="powo"/> {{div col|colwidth=20em}} *''[[Combretum aculeatum]]'' {{small|Vent.}} *''[[Combretum acuminatum]]'' {{small|Roxb.}} *''[[Combretum acutifolium]]'' {{small|Exell}} *''[[Combretum acutum]]'' {{small|M.A.Lawson}} *''[[Combretum adenogonium]]'' {{small|Steud. ex A.Rich.}} *''[[Combretum adrianii]]'' {{small|Jongkind}} *''[[Combretum afzelii]]'' {{small|G.Don}} *''[[Combretum alatum]]'' {{small|Craib}} *''[[Combretum albiflorum]]'' {{small|(Tul.) Jongkind}} *''[[Combretum albopunctatum]]'' {{small|Suess.}} *''[[Combretum alfredi]]'' {{small|Hance}} *''[[Combretum andongense]]'' {{small|Engl. & Diels}} *''[[Combretum andradae]]'' {{small|Exell & J.G.García}} *''[[Combretum angolense]]'' {{small|Welw. ex M.A.Lawson}} *''[[Combretum angustipetalum]]'' {{small|Chiov.}} *''[[Combretum annulatum]]'' {{small|Craib}} *''[[Combretum apetalum]]'' {{small|Wall. ex Kurz}} *''[[Combretum aphanopetalum]]'' {{small|Engl. & Diels}} *''[[Combretum apiculatum]]'' {{small|Sond.}} *''[[Combretum argenteum]]'' {{small|Bertol.}} *''[[Combretum argyrotrichum]]'' {{small|Welw. ex M.A.Lawson}} *''[[Combretum assimile]]'' {{small|Eichler}} *''[[Combretum atropurpureum]]'' {{small|Engl. & Diels}} *''[[Combretum aureonitens]]'' {{small|Engl. & Gilg}} *''[[Combretum auriculatum]]'' {{small|Engl. & Diels}} *''[[Combretum barbatum]]'' {{small|G.Don}} *''[[Combretum batesii]]'' {{small|Exell}} *''[[Combretum bauchiense]]'' {{small|Hutch. & Dalziel}} *''[[Combretum baumii]]'' {{small|Engl. & Gilg}} *''[[Combretum bipindense]]'' {{small|Engl. & Diels}} *''[[Combretum blepharopetalum]]'' {{small|Wickens}} *''[[Combretum boinensis]]'' {{small|Jongkind}} *''[[Combretum bracteatum]]'' {{small|(M.A.Lawson) Engl. & Diels}} *''[[Combretum bracteosum]]'' {{small|(Hochst.) Engl. & Diels}} *''[[Combretum brassiciforme]]'' {{small|Exell}} *''[[Combretum brevistylum]]'' {{small|Eichler}} *''[[Combretum brunneum]]'' {{small|Engl. & Diels}} *''[[Combretum butyrosum]]'' {{small|(G.Bertol.) Tul.}} *''[[Combretum cacoucia]]'' {{small|Exell ex Sandwith}} *''[[Combretum afrum]]'' {{small|(Eckl. & Zeyh.) Kuntze}} *''[[Combretum camporum]]'' {{small|Engl.}} *''[[Combretum capitatum]]'' {{small|De Wild. & Exell}} *''[[Combretum capituliflorum]]'' {{small|Fenzl ex Schweinf.}} *''[[Combretum capuronii]]'' {{small|Jongkind}} *''[[Combretum carringtonianum]]'' {{small|Exell & J.G.García}} *''[[Combretum caudatisepalum]]'' {{small|Exell & J.G.García}} *''[[Combretum celastroides]]'' {{small|Welw. ex M.A.Lawson}} *''[[Combretum chinense]]'' {{small|G.Don}} *''[[Combretum chionanthoides]]'' {{small|Engl. & Diels}} *''[[Combretum cinereopetalum]]'' {{small|Engl. & Diels}} *''[[Combretum cinnabarinum]]'' {{small|Engl. & Diels}} *''[[Combretum clarense]]'' {{small|Jongkind}} *''[[Combretum coccineum]]'' {{small|(Sonn.) Lam.}} *''[[Combretum collinum]]'' {{small|Fresen.}} *''[[Combretum comosum]]'' {{small|G.Don}} *''[[Combretum conchipetalum]]'' {{small|Engl. & Diels}} *''[[Combretum confertum]]'' {{small|(Benth.) M.A.Lawson}} *''[[Combretum × confusum]]'' {{small|Merr. & Rolfe}} *''[[Combretum congolanum]]'' {{small|Liben}} *''[[Combretum constrictum]]'' {{small|(Benth.) M.A.Lawson}} *''[[Combretum contractum]]'' {{small|Engl. & Diels}} *''[[Combretum coriifolium]]'' {{small|Engl. & Diels}} *''[[Combretum coursianum]]'' {{small|(H.Perrier) Jongkind}} *''[[Combretum cuspidatum]]'' {{small|Planch. ex Benth.}} *''[[Combretum decandrum]]'' {{small|Jacq.}} *''[[Combretum decaryi]]'' {{small|Jongkind}} *''[[Combretum deciduum]]'' {{small|Collett & Hemsl.}} *''[[Combretum demeusei]]'' {{small|De Wild.}} *''[[Combretum discolor]]'' {{small|Taub.}} *''[[Combretum dolichopodum]]'' {{small|Gilg}} *''[[Combretum duarteanum]]'' {{small|Cambess.}} *''[[Combretum echirense]]'' {{small|Jongkind}} *''[[Combretum edwardsii]]'' {{small|Exell}} *''[[Combretum elaeagnoides]]'' {{small|Klotzsch}} *''[[Combretum engleri]]'' {{small|Schinz, De Wild. & T.Durand}} *''[[Combretum eriogynum]]'' {{small|(H.Perrier) Jongkind}} *''[[Combretum erosum]]'' {{small|Jongkind}} *''[[Combretum erythrophloeum]]'' {{small|Gilg & Ledermann ex Engl.}} *''[[Combretum erythrophyllum]]'' {{small|(Burch.) Sond.}} *''[[Combretum esteriense]]'' {{small|Jongkind}} *''[[Combretum evisceratum]]'' {{small|(H.Perrier) Jongkind}} *''[[Combretum exalatum]]'' {{small|Engl.}} *''[[Combretum exannulatum]]'' {{small|(O.Hoffm.) Engl. & Diels}} *''[[Combretum excelsum]]'' {{small|Keay}} *''[[Combretum exellii]]'' {{small|Jongkind}} *''[[Combretum falcatum]]'' {{small|(Welw. ex Hiern) Jongkind}} *''[[Combretum farinosum]]'' {{small|Kunth}} *''[[Combretum flammeum]]'' {{small|Welw. ex Hiern}} *''[[Combretum foliatum]]'' {{small|Craib}} *''[[Combretum frangulifolium]]'' {{small|Kunth}} *''[[Combretum fruticosum]]'' {{small|(Loefl.) Stuntz}} *''[[Combretum fulvum]]'' {{small|Keay}} *''[[Combretum fuscum]]'' {{small|Planch. ex Benth.}} *''[[Combretum gabonense]]'' {{small|Exell}} *''[[Combretum germainii]]'' {{small|Liben}} *''[[Combretum ghesquierei]]'' {{small|Liben}} *''[[Combretum gillettianum]]'' {{small|Liben}} *''[[Combretum glabrum]]'' {{small|DC.}} *''[[Combretum glaucocarpum]]'' {{small|Mart.}} *''[[Combretum glutinosum]]'' {{small|Perr. ex DC.}} *''[[Combretum goetzei]]'' {{small|Engl. & Diels}} *''[[Combretum goldieanum]]'' {{small|F.Muell.}} *''[[Combretum goossensii]]'' {{small|De Wild. & Exell}} *''[[Combretum gordonii]]'' {{small|Jongkind}} *''[[Combretum gossweileri]]'' {{small|Exell}} *''[[Combretum gracile]]'' {{small|Schott}} *''[[Combretum graciliflorum]]'' {{small|Stace}} *''[[Combretum grandidieri]]'' {{small|Drake}} *''[[Combretum grandiflorum]]'' {{small|G.Don}} *''[[Combretum griffithii]]'' {{small|Van Heurck & Müll.Arg.}} *''[[Combretum harmsianum]]'' {{small|Diels}} *''[[Combretum harrisii]]'' {{small|Wickens}} *''[[Combretum hartmannianum]]'' {{small|Schweinf.}} *''[[Combretum haullevilleanum]]'' {{small|De Wild.}} *''[[Combretum hensii]]'' {{small|Engl. & Diels}} *''[[Combretum hereroense]]'' {{small|Schinz}} *''[[Combretum hilarianum]]'' {{small|D.Dietr.}} *''[[Combretum holstii]]'' {{small|Engl.}} *''[[Combretum homalioides]]'' {{small|Hutch. & Dalziel}} *''[[Combretum igneiflorum]]'' {{small|Rendón & R.Delgad.}} *''[[Combretum illairii]]'' {{small|Engl.}} *''[[Combretum imberbe]]'' {{small|Wawra}} *''[[Combretum indicum]]'' {{small|(L.) DeFilipps}} *''[[Combretum inflatum]]'' {{small|Jongkind}} *''[[Combretum ivanii]]'' {{small|Jongkind}} *''[[Combretum kasaiense]]'' {{small|Liben}} *''[[Combretum kirkii]]'' {{small|M.A.Lawson}} *''[[Combretum klossii]]'' {{small|Ridl.}} *''[[Combretum klotzschii]]'' {{small|Welw. ex M.A.Lawson}} *''[[Combretum kostermansii]]'' {{small|Exell}} *''[[Combretum kraussii]]'' {{small|Hochst.}} *''[[Combretum lanceolatum]]'' {{small|Pohl ex Eichler}} *''[[Combretum lanuginosum]]'' {{small|G.Don}} *''[[Combretum latialatum]]'' {{small|Engl.}} *''[[Combretum latifolium]]'' {{small|Blume}} *''[[Combretum laxum]]'' {{small|Jacq.}} *''[[Combretum lecardii]]'' {{small|Engl. & Diels}} *''[[Combretum leprosum]]'' {{small|Mart.}} *''[[Combretum lindense]]'' {{small|Exell & Mildbr.}} *''[[Combretum lisowskii]]'' {{small|Jongkind}} *''[[Combretum littoreum]]'' {{small|(Engl.) Engl.}} *''[[Combretum llewelynii]]'' {{small|J.F.Macbr.}} *''[[Combretum lokele]]'' {{small|Liben}} *''[[Combretum longicollum]]'' {{small|Jongkind}} *''[[Combretum longipilosum]]'' {{small|Engl. & Diels}} *''[[Combretum longispicatum]]'' {{small|(Engl.) Engl. & Diels}} *''[[Combretum longistipitatum]]'' {{small|Jongkind}} *''[[Combretum louisii]]'' {{small|Liben}} *''[[Combretum lukafuensis]]'' {{small|De Wild.}} *''[[Combretum luxenii]]'' {{small|Exell}} *''[[Combretum macrocalyx]]'' {{small|(Tul.) Jongkind}} *''[[Combretum malabaricum]]'' {{small|(Bedd.) Sujana, Ratheesh & Anil Kumar}} *''[[Combretum mannii]]'' {{small|G.Lawson ex Engl. & Diels}} *''[[Combretum marginatum]]'' {{small|Engl. & Diels}} *''[[Combretum marquesii]]'' {{small|Engl. & Diels}} *''[[Combretum mellifluum]]'' {{small|Eichler}} *''[[Combretum meridionalis]]'' {{small|(H.Perrier) Jongkind}} *''[[Combretum micranthum]]'' {{small|G.Don}} *''[[Combretum microphyllum]]'' {{small|Klotzsch}} *''[[Combretum mkuzense]]'' {{small|J.D.Carr & Retief}} *''[[Combretum moggii]]'' {{small|Exell}} *''[[Combretum molle]]'' {{small|R.Br. ex G.Don}} *''[[Combretum monetaria]]'' {{small|Mart.}} *''[[Combretum mooreanum]]'' {{small|Exell}} *''[[Combretum mortehanii]]'' {{small|De Wild. & Exell}} *''[[Combretum mossambicense]]'' {{small|(Klotzsch) Engl.}} *''[[Combretum mucronatum]]'' {{small|Schumach. & Thonn.}} *''[[Combretum multinervium]]'' {{small|Exell}} *''[[Combretum mussaendiflorum]]'' {{small|Engl. & Diels}} *''[[Combretum mweroense]]'' {{small|Baker}} *''[[Combretum nanum]]'' {{small|Buch.-Ham. ex D.Don}} *''[[Combretum ndjoleense]]'' {{small|Jongkind}} *''[[Combretum nigrescens]]'' {{small|King}} *''[[Combretum nigricans]]'' {{small|Lepr. ex Guill. & Perr.}} *''[[Combretum nioroense]]'' {{small|Aubrév. ex Keay}} *''[[Combretum niphophilum]]'' {{small|Gilg & Ledermann ex Engl.}} *''[[Combretum nusbaumeri]]'' {{small|Jongkind & L.Gaut.}} *''[[Combretum oatesii]]'' {{small|Rolfe}} *''[[Combretum obovatum]]'' {{small|F.Hoffm.}} *''[[Combretum obscurum]]'' {{small|Tul.}} *''[[Combretum octagonum]]'' {{small|(H.Perrier) Jongkind}} *''[[Combretum olivaceum]]'' {{small|Engl.}} *''[[Combretum oudenhovenii]]'' {{small|Jongkind}} *''[[Combretum ovalifolium]]'' {{small|Roxb. ex G.Don}} *''[[Combretum oxygonium]]'' {{small|(Tul.) Jongkind}} *''[[Combretum oxystachyum]]'' {{small|Welw. ex M.A.Lawson}} *''[[Combretum oyemense]]'' {{small|Exell}} *''[[Combretum padoides]]'' {{small|Engl. & Diels}} *''[[Combretum paniculatum]]'' {{small|Vent.}} *''[[Combretum paradoxum]]'' {{small|Welw. ex M.A.Lawson}} *''[[Combretum paraguariense]]'' {{small|(Eichler) Stace}} *''[[Combretum patelliforme]]'' {{small|Engl. & Diels}} *''[[Combretum paucinervium]]'' {{small|Engl. & Diels}} *''[[Combretum pavonii]]'' {{small|G.Don}} *''[[Combretum pecoense]]'' {{small|Exell}} *''[[Combretum pellegrinianum]]'' {{small|Exell}} *''[[Combretum pentagonum]]'' {{small|M.A.Lawson}} *''[[Combretum petrophilum]]'' {{small|Retief}} *''[[Combretum phaeocarpum]]'' {{small|Mart.}} *''[[Combretum pilosum]]'' {{small|Roxb. ex G.Don}} *''[[Combretum pisoniiflorum]]'' {{small|(Klotzsch) Engl.}} *''[[Combretum pisonioides]]'' {{small|Taub.}} *''[[Combretum platypterum]]'' {{small|(Welw.) Hutch. & Dalziel}} *''[[Combretum polyanthum]]'' {{small|Jongkind}} *''[[Combretum polystictum]]'' {{small|Hiern}} *''[[Combretum procursum]]'' {{small|Craib}} *''[[Combretum psidioides]]'' {{small|Welw.}} *''[[Combretum punctatum]]'' {{small|Blume}} *''[[Combretum purpureiflorum]]'' {{small|Engl.}} *''[[Combretum pyramidatum]]'' {{small|Desv.}} *''[[Combretum pyrifolium]]'' {{small|(C.Presl) Kurz}} *''[[Combretum quadrangulare]]'' {{small|Kurz}} *''[[Combretum quadratum]]'' {{small|Craib}} *''[[Combretum rabiense]]'' {{small|Jongkind}} *''[[Combretum racemosum]]'' {{small|P.Beauv.}} *''[[Combretum razianum]]'' {{small|K.G.Bhat}} *''[[Combretum recurvatum]]'' {{small|Sujana, Ratheesh & Anil Kumar}} *''[[Combretum relictum]]'' {{small|Hutch. & Dalziel}} *''[[Combretum rhodanthum]]'' {{small|Engl. & Diels}} *''[[Combretum riggenbachianum]]'' {{small|Gilg & Ledermann ex Engl.}} *''[[Combretum robustum]]'' {{small|Jongkind}} *''[[Combretum robynsii]]'' {{small|Exell}} *''[[Combretum rochetianum]]'' {{small|A.Rich. ex A.Juss.}} *''[[Combretum rohrii]]'' {{small|Exell}} *''[[Combretum rotundifolium]]'' {{small|Rich.}} *''[[Combretum rovirosae]]'' {{small|Exell}} *''[[Combretum roxburghii]]'' {{small|Spreng.}} *''[[Combretum rupestre]]'' {{small|Jongkind & Texier}} *''[[Combretum rupicola]]'' {{small|Ridl.}} *''[[Combretum sanjappae]]'' {{small|Chakrab. & Lakra}} *''[[Combretum scandens]]'' {{small|Liben}} *''[[Combretum schumannii]]'' {{small|Engl.}} *''[[Combretum sericeum]]'' {{small|G.Don}} *''[[Combretum shivannae]]'' {{small|Gholave, Kambale, Lekhak & S.R.Yadav}} *''[[Combretum sordidum]]'' {{small|Exell}} *''[[Combretum sphaeroides]]'' {{small|(Tul.) Jongkind}} *''[[Combretum spinosum]]'' {{small|Bonpl.}} *''[[Combretum stenopterum]]'' {{small|Exell}} *''[[Combretum stocksii]]'' {{small|Sprague}} *''[[Combretum struempellianum]]'' {{small|Gilg & Ledermann ex Engl.}} *''[[Combretum stylesii]]'' {{small|O.Maurin, Jordaan & A.E.van Wyk}} *''[[Combretum subglabratum]]'' {{small|De Wild.}} *''[[Combretum subumbellatum]]'' {{small|(Baker) Jongkind}} *''[[Combretum sundaicum]]'' {{small|Miq.}} *''[[Combretum sylvicola]]'' {{small|O.Maurin}} *''[[Combretum tarquense]]'' {{small|Clark}} *''[[Combretum tenuipetiolatum]]'' {{small|Wickens}} *''[[Combretum tessmannii]]'' {{small|Gilg ex Engl.}} *''[[Combretum tetragonocarpum]]'' {{small|Kurz}} *''[[Combretum tetralophoides]]'' {{small|Slooten}} *''[[Combretum tetralophum]]'' {{small|C.B.Clarke}} *''[[Combretum teuschii]]'' {{small|O.Hoffm.}} *''[[Combretum tibatiense]]'' {{small|Gilg & Ledermann ex Engl.}} *''[[Combretum tomentosum]]'' {{small|G.Don}} *''[[Combretum towaense]]'' {{small|Engl. & Diels}} *''[[Combretum trichophyllum]]'' {{small|Baker}} *''[[Combretum trifoliatum]]'' {{small|Vent.}} *''[[Combretum ulei]]'' {{small|Exell}} *''[[Combretum umbricola]]'' {{small|Engl.}} *''[[Combretum vendae]]'' {{small|A.E.van Wyk}} *''[[Combretum vernicosum]]'' {{small|Rusby}} *''[[Combretum villosum]]'' {{small|(Tul.) Jongkind}} *''[[Combretum violaceum]]'' {{small|(Tul.) Jongkind}} *''[[Combretum virgatum]]'' {{small|Welw. ex M.A.Lawson}} *''[[Combretum viscosum]]'' {{small|Exell}} *''[[Combretum wallichii]]'' {{small|DC.}} *''[[Combretum wattii]]'' {{small|Exell}} *''[[Combretum wilksii]]'' {{small|Jongkind}} *''[[Combretum winitii]]'' {{small|Craib}} *''[[Combretum xanthothyrsum]]'' {{small|Engl. & Diels}} *''[[Combretum youngii]]'' {{small|Exell}} *''[[Combretum zenkeri]]'' {{small|Engl. & Diels}} *''[[Combretum zeyheri]]'' {{small|Sond.}} {{div col end}} ==References== {{Reflist}} ==External links== {{Commons category}} {{AfricanPlants|Combretum}} {{Taxonbar|from=Q1777188}} [[Category:Combretum| ]] [[Category:Medicinal plants]] [[Category:Myrtales genera]]
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