Editing Cucujoidea

{{Short description|Superfamily of beetles}}
{{Automatic taxobox
| oldest_fossil = Jurassic
| image = Reitter-1911-plate098.png
| image_caption = Various Cucujoidea with larvae<br/>and anatomical details
| taxon = Cucujoidea
| display_parents = 3
| authority = [[Pierre André Latreille|Latreille]], 1802
}}

'''Cucujoidea''' is a [[Taxonomic rank|superfamily]] of [[beetles]]. This group formerly included all of the families now included in the superfamily [[Coccinelloidea]].<ref name=":0">{{Cite journal |last1=Robertson |first1=James A. |last2=Ślipiński |first2=Adam |last3=Moulton |first3=Matthew |last4=Shockley |first4=Floyd W. |last5=Giorgi |first5=Adriano |last6=Lord |first6=Nathan P. |last7=Mckenna |first7=Duane D. |last8=Tomaszewska |first8=Wioletta |last9=Forrester |first9=Juanita |last10=Miller |first10=Kelly B. |last11=Whiting |first11=Michael F. |year=2015 |title=Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia): Systematics of Cucujoidea and Coccinelloidea |url=https://onlinelibrary.wiley.com/doi/10.1111/syen.12138 |journal=Systematic Entomology |language=en |volume=40 |issue=4 |pages=745–778 |doi=10.1111/syen.12138|s2cid=55206626 |url-access=subscription }}</ref> They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark beetles ([[Scolytinae]]), which are [[weevil]]s (superfamily [[Curculionoidea]]).

== Morphology ==
The morphology of Cucujoidea is varied and there are no features uniting all members of the superfamily. In terms of general appearance, they tend to be small, drab in colour and with clubbed antennae.<ref name=":0" /> Even this is not universal; for example, ''[[Glischrochilus]]'' ([[Nitidulidae]]) have [[Aposematism|aposematic]] orange spots on their elytra.<ref>{{Cite journal |last=Clayhills |first=Tom |last2=Audisio |first2=Paolo |last3=Cline |first3=Andrew R. |last4=Mancini |first4=Emiliano |last5=Trizzino |first5=Marco |last6=Sabatelli |first6=Simone |date=2016-05-11 |title=Unraveling cryptic species diversity in an aposematic sap beetle genus (Coleoptera: Nitidulidae: Cryptarchinae) from northern Europe |url=https://www.researchgate.net/publication/292592741_Unraveling_cryptic_species_diversity_in_an_aposematic_sap_beetle_genus_Coleoptera_Nitidulidae_Cryptarchinae_from_northern_Europe |journal=Insect Systematics & Evolution |volume=47 |issue=2 |pages=131–148 |doi=10.1163/1876312X-47022137 |issn=1399-560X}}</ref>

Adults can be recognised by the procoxal cavities being internally open in most taxa, females having tarsal formula 5-5-5 and males 5-5-5 or 5-5-4 (rarely 4-4-4), females with tergite VIII concealed dorsally by tergite VII, and males with tergite X completely membraneous.<ref name=":0" />

Larvae have frontal arms usually lyriform, the mandible mesal surface usually with well-developed mola, a maxillary articulating area usually present, a hypopharyngeal sclerome usually present, and two pretarsal setae.<ref name=":0" />

== Ecology ==
Cucujoidea usually have cryptic habits, living in fungi, leaf litter or dead wood.<ref name=":0" /> This is reflected in many families having "fungus" or "bark" in their common names. The [[Kateretidae]]<ref>{{Cite web |title=Family Kateretidae - Short-winged Flower Beetles |url=https://bugguide.net/node/view/14000 |access-date=2023-08-25 |website=bugguide.net}}</ref> and some [[Phalacridae]]<ref>{{Cite web |title=Family Phalacridae - Shining Flower Beetles |url=https://bugguide.net/node/view/38232 |access-date=2023-08-25 |website=bugguide.net}}</ref> feed on flowers instead. The Nitidulidae are quite varied: some are [[Scavenger|saprophagous]] and [[Fungivore|mycetophagous]] like typical cucujoids, but others are associated with carrion, flowers, insect nests or stored food products.<ref>{{Cite web |title=Family Nitidulidae - Sap-feeding Beetles |url=https://bugguide.net/node/view/8441 |access-date=2023-08-25 |website=bugguide.net}}</ref>

==Taxonomy==
According to a 2015 revision, the following 25 families make up superfamily Cucujoidea:<ref name=":0" />
* [[Agapythidae]] <small>Sen Gupta and Crowson, 1969</small>
* [[Boganiidae]] <small>Sen Gupta & Crowson, 1966</small>
* [[Cavognathidae]] <small>Sen Gupta & Crowson, 1966</small>
* [[Cryptophagidae]] <small>Kirby, 1826</small> &ndash; [[silken fungus beetle]]s
* [[Cucujidae]] <small>Latreille, 1802</small> &ndash; [[flat bark beetle]]s
* [[Cybocephalidae]] <small>Jacquelin du Val, 1858</small>
* [[Cyclaxyridae]] <small>Gimmel, Leschen & Ślipiński, 2009</small> &ndash; sooty mould beetles
* [[Erotylidae]] <small>Latreille, 1802</small> &ndash; [[pleasing fungus beetle]]s
* [[Helotidae]] <small>Reitter, 1876</small>
* [[Hobartiidae]] <small>Sen Gupta & Crowson, 1966</small>
* [[Kateretidae]] <small>Erichson in Agassiz, 1846</small> &ndash; [[short-winged flower beetle]]s (= Brachypteridae)<!-- ICZN Op. 1916, 1999 -->
* [[Laemophloeidae]] <small>Ganglbauer 1899</small> &ndash; [[lined flat bark beetle]]s
* [[Lamingtoniidae]] <small>Sen Gupta & Crowson, 1966</small>
* [[Monotomidae]] <small>Laporte, 1840</small> &ndash; [[root-eating beetle]]s
* [[Myraboliidae]] <small>Lawrence and Britton, 1991</small>
* [[Nitidulidae]] <small>Latreille, 1802</small> &ndash; [[sap beetle]]s
* [[Passandridae]] <small>Erichson, 1845</small> &ndash; [[parasitic flat bark beetle]]s
* [[Phalacridae]] <small>Leach, 1815</small> &ndash; [[shining flower beetle]]s
* [[Phloeostichidae]] <small>Reitter, 1911</small>
* [[Priasilphidae]] <small>Crowson, 1973</small>
* [[Protocucujidae]] <small>Crowson, 1954</small>
* [[Silvanidae]] <small>Kirby, 1937</small> &ndash; [[silvanid flat bark beetle]]s
* [[Smicripidae]] <small>Horn, 1879</small> &ndash; [[palmetto beetle]]s
* [[Sphindidae]] <small>Jacquelin du Val, 1860</small> &ndash; [[dry-fungus beetle]]s
* [[Tasmosalpingidae]] <small>Lawrence and Britton, 1991</small>
* {{Extinct}}[[Parandrexidae]] <small>Kirejtshuk, 1994</small> (Middle Jurassic-Early Cretaceous)
* {{Extinct}}[[Wabbelidae]] <small>Alekseev, 2017</small><ref>{{cite journal |author=Vitalii I. Alekseev |year=2017 |title=A new coleopterous family Wabbelidae fam. nov. (Coleoptera: Cucujoidea) from Baltic amber (Cenozoic, Paleogene, Eocene) |journal=Baltic Journal of Coleopterology |volume=17 |issue=1 |pages=29–41 |url=http://www.bjc.sggw.pl/viewart.php?bookid=34&artid=578 }}</ref> (Eocene)

=== Extinct genera ===

* {{extinct}}''[[Alloterocucus]]'' <small>Li et al., 2022</small> [[Burmese amber]], Myanmar, mid-[[Cretaceous]] (latest [[Albian]]- earliest [[Cenomanian]])

==References==
{{Reflist}}

==External links==
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[[Category:Cucujoidea| ]]
[[Category:Beetle superfamilies]]
[[Category:Taxa named by Pierre André Latreille]]