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Gadiformes
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{{Short description|Order of fishes}} {{Automatic taxobox | fossil_range = {{fossil range|Maastrichtian|present}} | image = Atlantic-cod-1.jpg | image_upright = 1.15 | image_caption = ''[[Gadus morhua]]'' | taxon = Gadiformes | authority = [[Edwin Stephen Goodrich|Goodrich]], 1909 | subdivision_ranks = Families | subdivision = See text | type_species = ''[[Gadus morhua]]'' | type_species_authority = Linnaeus 1758 }} '''Gadiformes''' {{IPAc-en|'|g|æ|d|ᵻ|f|ɔːr|m|iː|z}}, also called the '''Anacanthini''', are an [[order (biology)|order]] of [[ray-finned fish]] that include the [[cod]], [[hake]]s, [[pollock]], [[haddock]], [[burbot]], [[Lotidae|rockling]]s and [[Moridae|mora]]s, many of which are [[food fish]] of major commercial value. They are mostly [[marine fish]] found throughout the world and the vast majority are found in [[temperate]] or colder regions (tropical species are typically deep-water) while a few species may enter [[brackish]] [[estuaries]]. [[Microgadus proximus|Pacific tomcod]]s, one of the two species that makes up the genus ''[[Microgadus]]'', are able to enter freshwater, but there is no evidence that they breed there. Some populations of landlocked [[Microgadus tomcod|Atlantic tomcod]] on the other hand, complete their entire life cycle in freshwater. Yet only one species, the [[burbot]] (''Lota lota''), is a true [[freshwater fish]].<ref>[https://books.google.com/books?id=UO6nAgAAQBAJ&dq=Gadidae+tomcod+landlocked+life+cycle+burbot&pg=PA417 The Freshwater Fishes of British Columbia]</ref> Common characteristics include the positioning of the [[pelvic fin]]s (if present), below or in front of the [[pectoral fin]]s. Gadiformes are [[physoclist]]s, which means their [[swim bladder]]s do not have a [[pneumatic duct]]. The fins are spineless. Gadiform fish range in size from the [[codlet]]s, which may be as small as {{convert|7|cm|in|abbr=on}} in adult length, to the [[Atlantic cod]], ''Gadus morhua'', which reaches up to {{convert|2|m|ft|abbr=on}}.<ref name=EoF>{{cite book |editor=Paxton, J.R. |editor2=Eschmeyer, W.N.|author= Cohen, D.M.|year=1998|title=Encyclopedia of Fishes|publisher= Academic Press|location=San Diego|pages= 130–132|isbn= 0-12-547665-5}}</ref> The earliest gadiforms are ''[[Palaeogadus|Palaeogadus weltoni]]'' from the [[Maastrichtian]] of the United States and the undescribed, informally named "''Protocodus''" from the [[Danian|Early Paleocene]] of [[Greenland]].<ref>{{Cite journal |last1=Stringer |first1=Gary |last2=Schwarzhans |first2=Werner |date=2021-09-01 |title=Upper Cretaceous teleostean otoliths from the Severn Formation (Maastrichtian) of Maryland, USA, with an unusual occurrence of Siluriformes and Beryciformes and the oldest Atlantic coast Gadiformes |journal=Cretaceous Research |volume=125 |pages=104867 |doi=10.1016/j.cretres.2021.104867 |issn=0195-6671|doi-access=free |bibcode=2021CrRes.12504867S }}</ref><ref>{{Cite journal |last1=Friedman |first1=Matt |last2=V. Andrews |first2=James |last3=Saad |first3=Hadeel |last4=El-Sayed |first4=Sanaa |date=2023-06-16 |title=The Cretaceous–Paleogene transition in spiny-rayed fishes: surveying "Patterson's Gap" in the acanthomorph skeletal record André Dumont medalist lecture 2018 |url=https://popups.uliege.be/1374-8505/index.php?id=7048 |journal=Geologica Belgica |language=en |doi=10.20341/gb.2023.002 |issn=1374-8505|doi-access=free }}</ref> == Taxonomy == The following classification is based on ''[[Eschmeyer's Catalog of Fishes]]'':<ref name=":13">{{Cite web |last=Fricke |first=R. |last2=Eschmeyer |first2=W. N. |last3=Van der Laan |first3=R. |date=2025 |title=ESCHMEYER'S CATALOG OF FISHES: CLASSIFICATION |url=https://www.calacademy.org/eschmeyers-catalog-of-fishes-classification |access-date=2025-02-10 |website=California Academy of Sciences |language=en}}</ref> * Order '''Gadiformes''' ** Suborder Stylephoroidei *** Family [[Stylephoridae]] <small>[[William Swainson|Swainson]], 1839</small> (tube-eyes or threadtails) ** Suborder Bregmacerotoidei *** Family [[Codlet|Bregmacerotidae]] <small>[[Theodore Gill|Gill]], 1872</small> (codlets) ** Suborder Gadoidei *** Family [[Phycidae]] <small>Swainson, 1838</small> (phycid hakes) *** Family [[Gaidropsaridae]] <small>[[David Starr Jordan|Jordan]] & [[Barton Warren Evermann|Evermann]], 1898</small> (rocklings) *** Family [[Lotidae]] <small>[[Charles Lucien Bonaparte|Bonaparte]], 1835</small> (hakes and burbots) *** Family [[Gadidae]] <small>[[Constantine Samuel Rafinesque|Rafinesque]], 1810</small> (cods and haddocks) ** Suborder Ranicipitoidei *** Family [[Ranicipitidae]] <small>Bonaparte, 1835</small> (tadpole fishes) ** Suborder Merluccioidei *** Family [[Merlucciidae]] <small>Rafinesque, 1815</small> (merlucciid hakes) ** Suborder Macrouroidei *** Family [[Euclichthyidae]] <small>Cohen, 1984</small> (Eucla cods) *** Family [[Muraenolepididae]] <small>[[Charles Tate Regan|Regan]], 1903</small> (eel cods) *** Family [[Melanonidae]] <small>[[George Brown Goode|Goode]] & [[Tarleton Hoffman Bean|Bean]], 1896</small> (arrowtails or pelagic cods) *** Family [[Trachyrincidae]] <small>Goode & Bean, 1896</small> (armoured grenadiers) *** Family [[Moridae]] <small>Moreau, 1881</small> (codlings or deepsea cods) *** Family [[Macruronidae]] <small>Regan, 1903</small> (blue grenadiers) *** Family [[Lyconidae]] <small>[[Albert Günther|Günther]], 1887</small> (Atlantic hakes) *** Family [[Bathygadidae]] <small>Jordan & Evermann, 1898</small> (rattails) *** Family [[Steindachneriidae]] <small>[[Albert Eide Parr|Parr]], 1942</small> (luminous hakes) *** Family [[Macrouridae]] <small>Bonaparte, 1831</small> (grenadiers or rattails)<ref name="Iwamoto2015">Iwamoto, T., Nakayama, N., Shao, K.-T. & Ho, H.-C. (2015): Synopsis of the Grenadier Fishes (Gadiformes; Teleostei) of Taiwan. ''Proceedings of the California Academy of Sciences, (Series 4), 62 (3): 31–126.''</ref> ==Timeline of genera== <timeline> ImageSize = width:900px height:auto barincrement:15px PlotArea = left:10px bottom:50px top:10px right:50px Period = from:-145.5 till:10 TimeAxis = orientation:horizontal ScaleMajor = unit:year increment:10 start:-145.5 ScaleMinor = unit:year increment:1 start:-145.5 TimeAxis = orientation:hor AlignBars = justify Colors = #legends id:CAR value:claret id:ANK value:rgb(0.4,0.3,0.196) id:HER value:teal id:HAD value:green id:OMN value:blue id:black value:black id:white value:white id:cretaceous value:rgb(0.5,0.78,0.31) id:earlycretaceous value:rgb(0.63,0.78,0.65) id:latecretaceous value:rgb(0.74,0.82,0.37) id:cenozoic value:rgb(0.54,0.54,0.258) id:paleogene value:rgb(0.99,0.6,0.32) id:paleocene value:rgb(0.99,0.65,0.37) id:eocene value:rgb(0.99,0.71,0.42) id:oligocene value:rgb(0.99,0.75,0.48) id:neogene value:rgb(0.999999,0.9,0.1) id:miocene value:rgb(0.999999,0.999999,0) id:pliocene value:rgb(0.97,0.98,0.68) id:quaternary value:rgb(0.98,0.98,0.5) id:pleistocene value:rgb(0.999999,0.95,0.68) id:holocene value:rgb(0.999,0.95,0.88) BarData= bar:eratop bar:space bar:periodtop bar:space bar:NAM1 bar:NAM2 bar:NAM3 bar:NAM4 bar:NAM5 bar:NAM6 bar:NAM7 bar:NAM8 bar:NAM9 bar:NAM10 bar:NAM11 bar:NAM12 bar:NAM13 bar:NAM14 bar:NAM15 bar:NAM16 bar:NAM17 bar:NAM18 bar:NAM19 bar:NAM20 bar:NAM21 bar:NAM22 bar:NAM23 bar:NAM24 bar:NAM25 bar:NAM26 bar:NAM27 bar:NAM28 bar:NAM29 bar:NAM30 bar:NAM31 bar:NAM32 bar:NAM33 bar:NAM34 bar:NAM35 bar:NAM36 bar:NAM37 bar:NAM38 bar:NAM39 bar:NAM40 bar:NAM41 bar:NAM42 bar:NAM43 bar:NAM44 bar:NAM45 bar:NAM46 bar:NAM47 bar:NAM48 bar:NAM49 bar:NAM50 bar:NAM51 bar:NAM52 bar:space bar:period bar:space bar:era PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 shift:(7,-4) bar:periodtop from: -145.5 till: -99.6 color:earlycretaceous text:[[Early Cretaceous|Early]] from: -99.6 till: -65.5 color:latecretaceous text:[[Late Cretaceous|Late]] from: -65.5 till: -55.8 color:paleocene text:[[Paleocene|Paleo.]] from: -55.8 till: -33.9 color:eocene text:[[Eocene|Eo.]] from: -33.9 till: -23.03 color:oligocene text:[[Oligocene|Oligo.]] from: -23.03 till: -5.332 color:miocene text:[[Miocene|Mio.]] from: -5.332 till: -2.588 color:pliocene text:[[Pliocene|Pl.]] from: -2.588 till: -0.0117 color:pleistocene text:[[Pleistocene|Pl.]] from: -0.0117 till: 0 color:holocene text:[[Holocene|H.]] bar:eratop from: -145.5 till: -65.5 color:cretaceous text:[[Cretaceous]] from: -65.5 till: -23.03 color:paleogene text:[[Paleogene]] from: -23.03 till: -2.588 color:neogene text:[[Neogene]] from: -2.588 till: 0 color:quaternary text:[[Quaternary|Q.]] PlotData= align:left fontsize:M mark:(line,white) width:5 anchor:till align:left color:latecretaceous bar:NAM1 from: -85.8 till: -70.6 text: [[Rankinian]] color:latecretaceous bar:NAM2 from: -83.5 till: -65.5 text: [[Paratichthys]] color:eocene bar:NAM3 from: -55.8 till: -48.6 text: [[Rhinocephalus]] color:eocene bar:NAM4 from: -55.8 till: -37.2 text: [[Trichurides]] color:eocene bar:NAM5 from: -55.8 till: -11.608 text: [[Palaeogadus]] color:eocene bar:NAM6 from: -55.8 till: 0 text: [[Urophycis]] color:eocene bar:NAM7 from: -55.8 till: 0 text: [[Raniceps]] color:eocene bar:NAM8 from: -55.8 till: 0 text: [[Nezumia]] color:eocene bar:NAM9 from: -55.8 till: 0 text: [[Merluccius]] color:eocene bar:NAM10 from: -55.8 till: 0 text: [[Melanonus]] color:eocene bar:NAM11 from: -48.6 till: 0 text: [[Bregmaceros]] color:eocene bar:NAM12 from: -37.2 till: 0 text: [[Tripterophycis]] color:eocene bar:NAM13 from: -37.2 till: 0 text: [[Macruronus]] color:eocene bar:NAM14 from: -37.2 till: 0 text: [[Eulichthys]] color:eocene bar:NAM15 from: -37.2 till: 0 text: [[Coelorhynchus]] color:eocene bar:NAM16 from: -37.2 till: 0 text: [[Bathygadus]] color:oligocene bar:NAM17 from: -33.9 till: -28.4 text: [[Eophycis]] color:oligocene bar:NAM18 from: -33.9 till: 0 text: [[Trachyrhinchus]] color:oligocene bar:NAM19 from: -33.9 till: 0 text: [[Squalogadus]] color:oligocene bar:NAM20 from: -33.9 till: 0 text: [[Phycis]] color:oligocene bar:NAM21 from: -33.9 till: 0 text: [[Lota]] color:oligocene bar:NAM22 from: -33.9 till: 0 text: [[Hymenocephalus]] color:oligocene bar:NAM23 from: -33.9 till: 0 text: [[Gaidropsarus]] color:oligocene bar:NAM24 from: -33.9 till: 0 text: [[Gadiculus]] color:oligocene bar:NAM25 from: -28.4 till: -15.97 text: [[Pseudoraniceps]] color:oligocene bar:NAM26 from: -28.4 till: -11.608 text: [[Palaeomolva]] color:oligocene bar:NAM27 from: -28.4 till: 0 text: [[Trisopterus]] color:oligocene bar:NAM28 from: -28.4 till: 0 text: [[Micromesistius]] color:oligocene bar:NAM29 from: -28.4 till: 0 text: [[Coryphaenoides]] color:oligocene bar:NAM30 from: -28.4 till: 0 text: [[Brosmius]] color:miocene bar:NAM31 from: -23.03 till: -15.97 text: [[Bregmacerinia]] color:miocene bar:NAM32 from: -23.03 till: -5.332 text: [[Trichiurichthys]] color:miocene bar:NAM33 from: -23.03 till: 0 text: [[Ventrifossa]] color:miocene bar:NAM34 from: -23.03 till: 0 text: [[Physiculus]] color:miocene bar:NAM35 from: -23.03 till: 0 text: [[Austrophycis]] color:miocene bar:NAM36 from: -15.97 till: 0 text: [[Molva]] color:miocene bar:NAM37 from: -15.97 till: 0 text: [[Merlangius]] color:miocene bar:NAM38 from: -15.97 till: 0 text: [[Lepidion]] color:miocene bar:NAM39 from: -15.97 till: 0 text: [[Laemonema]] color:miocene bar:NAM40 from: -11.608 till: -5.332 text: [[Eclipes]] color:miocene bar:NAM41 from: -11.608 till: -5.332 text: [[Bolbocara]] color:miocene bar:NAM42 from: -11.608 till: 0 text: [[Paratrisopterus]] color:miocene bar:NAM43 from: -11.608 till: 0 text: [[Gadus]] color:miocene bar:NAM44 from: -11.608 till: 0 text: [[Gadomus]] color:miocene bar:NAM45 from: -11.608 till: 0 text: [[Brosme]] color:pliocene bar:NAM46 from: -5.332 till: -2.588 text: [[Pseudophycis]] color:pliocene bar:NAM47 from: -5.332 till: -2.588 text: [[Gadella]] color:pliocene bar:NAM48 from: -5.332 till: 0 text: [[Theragra]] color:pliocene bar:NAM49 from: -5.332 till: 0 text: [[Microgadus]] color:pliocene bar:NAM50 from: -5.332 till: 0 text: [[Melanogrammus]] color:pliocene bar:NAM51 from: -5.332 till: 0 text: [[Lycodopsis]] color:pleistocene bar:NAM52 from: -2.588 till: -0.0117 text: [[Pollachius]] PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 bar:period from: -145.5 till: -99.6 color:earlycretaceous text:[[Early Cretaceous|Early]] from: -99.6 till: -65.5 color:latecretaceous text:[[Late Cretaceous|Late]] from: -65.5 till: -55.8 color:paleocene text:[[Paleocene|Paleo.]] from: -55.8 till: -33.9 color:eocene text:[[Eocene|Eo.]] from: -33.9 till: -23.03 color:oligocene text:[[Oligocene|Oligo.]] from: -23.03 till: -5.332 color:miocene text:[[Miocene|Mio.]] from: -5.332 till: -2.588 color:pliocene text:[[Pliocene|Pl.]] from: -2.588 till: -0.0117 color:pleistocene text:[[Pleistocene|Pl.]] from: -0.0117 till: 0 color:holocene text:[[Holocene|H.]] bar:era from: -145.5 till: -65.5 color:cretaceous text:[[Cretaceous]] from: -65.5 till: -23.03 color:paleogene text:[[Paleogene]] from: -23.03 till: -2.588 color:neogene text:[[Neogene]] from: -2.588 till: 0 color:quaternary text:[[Quaternary|Q.]] </timeline> ==References== {{Reflist}} {{Actinopterygii}} {{Taxonbar|from=Q216241}} {{Authority control}} [[Category:Gadiformes| ]] [[Category:Taxa named by Edwin Stephen Goodrich]] [[Category:Ray-finned fish orders]] [[Category:Extant Maastrichtian first appearances]]
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