Editing Lateral geniculate nucleus

{{short description|Component of the visual system in the brain's thalamus}}
{{Redirect|LGN}}
{{Infobox brain
| Name            = Lateral geniculate nucleus
| Latin           = corpus geniculatum laterale
| Image           = Gray719.png
| Caption         = [[Hindbrain|Hind-]] and [[mid-brain]]s; postero-lateral view. (Lateral geniculate body visible near top.)
| Image2          =
| Caption2        =
| IsPartOf        = [[Thalamus]]
| System          = [[Visual system|Visual]]
| Components      =
| Artery          = [[Anterior choroidal artery|Anterior choroidal]] and [[Posterior cerebral artery|Posterior cerebral]]
| Vein            = [[Terminal vein]]
| Acronym         = LGN
}}

In [[neuroanatomy]], the '''lateral geniculate nucleus''' ('''LGN'''; also called the '''lateral geniculate body''' or '''lateral geniculate complex''') is a structure in the [[thalamus]] and a key component of the mammalian [[visual pathway]]. It is a small, ovoid, [[Anatomical terms of location#Dorsal_and_ventral|ventral]] projection of the thalamus where the thalamus connects with the [[optic nerve]]. There are two LGNs, one on the left and another on the right side of the thalamus. In humans, both LGNs have six layers of [[neuron]]s ([[grey matter]]) alternating with optic fibers ([[white matter]]).

The LGN receives information directly from the ascending [[retinal ganglion cell]]s via the optic tract and from the [[reticular activating system]]. Neurons of the LGN send their axons through the [[optic radiation]], a direct pathway to the [[primary visual cortex]]. In addition, the LGN receives many strong feedback connections from the primary [[visual cortex]].<ref name=Cudeiro2006>{{cite journal|last=Cudeiro|first=Javier|author2=Sillito, Adam M. |title=Looking back: corticothalamic feedback and early visual processing|journal=Trends in Neurosciences|year=2006|pages=298–306|volume=29|issue=6|pmid=16712965|doi=10.1016/j.tins.2006.05.002|citeseerx=10.1.1.328.4248|s2cid=6301290}}</ref> In [[humans]] as well as other [[mammals]], the two strongest pathways linking the eye to the brain are those projecting to the dorsal part of the LGN in the thalamus, and to the [[superior colliculus]].<ref>Goodale, M. & Milner, D. (2004)''Sight unseen.''Oxford University Press, Inc.: New York.</ref>

==Structure==
[[File:Thalmus.png|thumb|240px|Nuclei of the Thalamus]]
Both the left and right [[Cerebral hemisphere|hemisphere]]s of the [[brain]] have a lateral geniculate nucleus, named after its resemblance to a bent knee (''genu'' is Latin for "knee"). In [[human]]s as well as in many other [[primate]]s, the LGN has layers of [[magnocellular cells]] and [[parvocellular cells]] that are interleaved with layers of koniocellular cells.

In humans the LGN is normally described as having six distinctive layers. The inner two layers, (1 and 2) are [[magnocellular cell|magnocellular layers]], while the outer four layers, (3, 4, 5 and 6), are [[parvocellular cell|parvocellular layers]].  An additional set of neurons, known as the [[koniocellular cell|koniocellular layers]], are found ventral to each of the magnocellular and parvocellular layers.<ref name=Brodal2010>{{cite book|last1=Brodal|first1=Per|title=The central nervous system : structure and function|date=2010|publisher=Oxford University Press|location=New York|isbn=978-0-19-538115-3|edition=4th}}</ref>{{rp|227ff}}<ref name="Carlson, N. R. 2007">{{cite book|last1=Carlson|first1=Neil R.|title=Physiology of behavior|date=2007|publisher=Pearson/Allyn & Bacon|location=Boston|isbn=978-0205467242|edition=9th}}</ref>  This layering is variable between primate species, and extra leafleting is variable within species.

The average volume of each LGN in an adult human is about 118mm<math>{}^3</math>. (This is the same volume as a 4.9mm-sided cube.) A study of 24 hemispheres from 15 normal individuals with average age 59 years at autopsy found variation from about 91 to 157mm<math>{}^3</math>.<ref>{{cite journal |last1=Andrews |first1=Timothy J. |last2=Halpern |first2=Scott D. |last3=Purves |first3=Dale |title=Correlated Size Variations in Human Visual Cortex, Lateral Geniculate Nucleus, and Optic Tract |journal=Journal of Neuroscience |date=1997 |volume=17 |issue=8 |pages=2859–2868 |doi=10.1523/JNEUROSCI.17-08-02859.1997 |doi-access=free|pmc=6573115 }}</ref> The same study found that in each LGN, the magnocellular layers comprised about 28mm<math>{}^3</math> in total, and the parvocellular layers comprised about 90mm<math>{}^3</math> in total.

==M, P, K cells==
[[File:Cgl Nissl2.svg|thumb|right|240px|Relative locations of the M-, P-, and K-layers (macaque monkey)]]

{| class="wikitable"
! Type || Size* || [[Retinal ganglion cell|RGC]] Source || Type of Information || Location || Response || Number
 |-
 | M: [[Magnocellular cells]] || Large || [[Parasol cell]]s || perception of movement, depth, and small differences in brightness || Layers 1 and 2  || rapid and transient || ?
 |-
 | P: [[Parvocellular cells]] (or "parvicellular")   || Small || [[Midget cell]]s || perception of color and form (fine details) || Layers 3, 4, 5 and 6 || slow and sustained || ?
 |- /
 | K: [[Koniocellular cells]] (or "interlaminar") ||  Very small cell bodies || [[Bistratified cell]]s || || Between each of the M and P layers || ||
|}
<nowiki>*</nowiki>Size describes the cell body and dendritic tree, though also can describe the receptive field

The magnocellular, parvocellular, and koniocellular layers of the LGN correspond with the similarly named types of [[retinal ganglion cell]]s. Retinal P ganglion cells send axons to a parvocellular layer, M ganglion cells send axons to a magnocellular layer, and K ganglion cells send axons to a koniocellular layer.<ref>{{cite book |last1=Purves|first1=Dale|last2=Augustine|first2=George|last3=Fitzpatrick|first3=David|last4=Hall|first4=William|last5=Lamantia|first5=Anthony-Samuel|last6=White|first6=Leonard|title=Neuroscience|date=2011|publisher=Sinauer|location=Sunderland, Mass.|isbn=978-0878936953|edition=5.}}</ref>{{rp|269}}

Koniocellular cells are functionally and neurochemically distinct from M and P cells and provide a third channel to the visual cortex. They project their axons between the layers of the lateral geniculate nucleus where M and P cells project. Their role in visual perception is presently unclear; however, the koniocellular system has been linked with the integration of somatosensory system-proprioceptive information with visual perception{{Citation needed|date=November 2023}}, and it may also be involved in color perception.<ref>{{cite journal|last1=White|first1=BJ|last2=Boehnke|first2=SE|last3=Marino|first3=RA|last4=Itti|first4=L|last5=Munoz|first5=DP|title=Color-related signals in the primate superior colliculus|journal=The Journal of Neuroscience|date=Sep 30, 2009|volume=29|issue=39|pages=12159–66|doi=10.1523/JNEUROSCI.1986-09.2009|pmid=19793973|pmc=6666157|doi-access=free}}</ref>

The parvo- and magnocellular fibers were previously thought to dominate the Ungerleider–Mishkin [[ventral stream]] and [[dorsal stream]], respectively. However, new evidence has accumulated showing that the two streams appear to feed on a more even mixture of different types of nerve fibers.<ref>Goodale & Milner, 1993, 1995.</ref>

The other major retino–cortical visual pathway is the [[tectopulvinar pathway]], routing primarily through the [[superior colliculus]] and thalamic [[pulvinar nuclei|pulvinar]] nucleus onto [[posterior parietal]] cortex and [[visual area MT]].

==Ipsilateral and contralateral layers==
Both the LGN in the right hemisphere and the LGN in the left hemisphere receive input from each eye. However, each LGN only receives information from one half of the visual field. [[Retinal ganglion cell]]s (RGCs) from the inner halves of each retina (the nasal sides) [[Decussation|decussate]] (cross to the other side of the brain) through the [[optic chiasma]] (''khiasma'' means "cross-shaped"). RGCs from the outer half of each retina (the temporal sides) remain on the same side of the brain. Therefore, the right LGN receives visual information from the left visual field, and the left LGN receives visual information from the right visual field. Within one LGN, the visual information is divided among the various layers as follows:<ref>Nicholls J., ''et al.'' ''From Neuron to Brain: Fourth Edition''. Sinauer Associates, Inc. 2001.</ref>
* the eye on the same side (the ''ipsilateral'' eye) sends information to layers 2, 3 and 5
* the eye on the opposite side (the ''contralateral'' eye) sends information to layers 1, 4 and 6.

This description applies to the LGN of many primates, but not all. The sequence of layers receiving information from the ipsilateral and contralateral (opposite side of the head) eyes is different in the [[tarsier]].<ref>{{cite journal|last1=Rosa|first1=MG|last2=Pettigrew|first2=JD|last3=Cooper|first3=HM|title=Unusual pattern of retinogeniculate projections in the controversial primate Tarsius|journal=Brain, Behavior and Evolution|date=1996|volume=48|issue=3|pages=121–9|pmid=8872317|doi=10.1159/000113191}}</ref> Some neuroscientists suggested that "this apparent difference distinguishes tarsiers from all other primates, reinforcing the view that they arose in an early, independent line of primate evolution".<ref>{{cite journal|last1=Collins|first1=CE|last2=Hendrickson|first2=A|last3=Kaas|first3=JH|title=Overview of the visual system of Tarsius|journal=The Anatomical Record Part A: Discoveries in Molecular, Cellular, and Evolutionary Biology|date=Nov 2005|volume=287|issue=1|pages=1013–25|pmid=16200648|doi=10.1002/ar.a.20263|doi-access=free}}</ref>

== Input ==
The LGN receives input from the retina and many other brain structures, especially visual cortex.

The principal neurons in the LGN receive strong inputs from the retina. However, the retina only accounts for a small percentage of LGN input. As much as 95% of input in the LGN comes from the visual cortex, superior colliculus, pretectum, thalamic reticular nuclei, and local LGN interneurons. Regions in the brainstem that are not involved in visual perception also project to the LGN, such as the mesencephalic reticular formation, dorsal raphe nucleus, periaqueuctal grey matter, and the locus coeruleus.<ref name="Guillery 163–75">{{cite journal|last=Guillery|first=R|author2=SM Sherman |title=Thalamic relay functions and their role in corticocortical communication: generalizations from the visual system|journal=Neuron|date=Jan 17, 2002|volume=33|issue=2|pages=163–75|doi=10.1016/s0896-6273(01)00582-7 |pmid=11804565|doi-access=free}}</ref>  The LGN also receives some inputs from the [[optic tectum]] (known as the [[superior colliculus]] in mammals).<ref>In Chapter 7, section "The Parcellation Hypothesis" of "Principles of Brain Evolution", [[Georg F. Striedter]] (Sinauer Associates, Sunderland, MA, USA, 2005) states, "...we now know that the LGN receives at least some inputs from the optic tectum (or superior colliculus) in many amniotes". He cites "{{cite journal | last1 = Wild | first1 = J.M. | year = 1989 | title = Pretectal and tectal projections to the homolog of the dorsal lateral geniculate nucleus in the pigeon—an anterograde and retrograde tracing study with cholera-toxin conjugated to horseradish-peroxidase | doi = 10.1016/0006-8993(89)91342-5 | journal = Brain Res | volume = 479 | issue = 1 | pages = 130–137 | pmid = 2924142 | s2cid = 29034684 }}" and also "Kaas, J.H., and Huerta, M.F. 1988. The subcortical visual system of primates. In: Steklis H. D., Erwin J., editors. Comparative primate biology, vol 4: neurosciences. New York: Alan Liss, pp. 327–391.</ref> These non-retinal inputs can be excitatory, inhibitory, or modulatory.<ref name="Guillery 163–75"/>

== Output ==
Information leaving the LGN travels out on the [[optic radiation]]s, which form part of the ''retrolenticular portion'' of the [[internal capsule]].

The [[axon]]s that leave the LGN go to V1 [[visual cortex]]. Both the magnocellular layers 1–2 and the parvocellular layers 3–6 send their axons to layer 4 in V1. Within layer 4 of V1, layer 4cβ receives parvocellular input, and layer 4cα receives magnocellular input. However, the koniocellular layers, intercalated between LGN layers 1–6 send their axons primarily to the cytochrome-oxidase rich blobs of layers 2 and 3 in V1.<ref name=HendryandReid2000>{{cite journal|last=Hendry|first=Stewart H. C.|author2=Reid, R. Clay|title=The koniocellular pathway in primate vision|journal=Annual Review of Neuroscience|year=2000|pmid=10845061|pages=127–153|volume=23|doi=10.1146/annurev.neuro.23.1.127}}</ref> [[Axon]]s from layer 6 of [[visual cortex]] send information back to the LGN.

Studies involving [[blindsight]] have suggested that projections from the LGN travel not only to the primary visual cortex but also to higher cortical areas V2 and V3. Patients with blindsight are phenomenally blind in certain areas of the visual field corresponding to a contralateral lesion in the primary visual cortex; however, these patients are able to perform certain motor tasks accurately in their blind field, such as grasping. This suggests that neurons travel from the LGN to both the primary visual cortex and higher cortex regions.<ref name=Schmid2010>{{cite journal|last=Schmid|first=Michael C.|author2=Mrowka, Sylwia W. |author3=Turchi, Janita|title=Blindsight depends on the lateral geniculate nucleus|journal=Nature|year=2010|doi=10.1038/nature09179 |pages=373–377|volume=466|issue=7304|display-authors=etal|pmid=20574422|pmc=2904843|bibcode=2010Natur.466..373S}}</ref>

==Function in visual perception==
The output of the LGN serves several functions.

Computations are achieved to determine the position of every major element in object space relative to the principal plane.  Through subsequent motion of the eyes, a larger stereoscopic mapping of the visual field is achieved.<ref>Lindstrom, S. & Wrobel, A. (1990) Intracellular recordings from binocularly activated cells in the cats dorsal lateral geniculate nucleus Acta Neurobiol Exp vol 50, pp 61–70</ref>

It has been shown that while the retina accomplishes spatial [[decorrelation]] through center surround inhibition, the LGN accomplishes temporal decorrelation.<ref>Dawei W. Dong and Joseph J. Atick, Network–Temporal Decorrelation: A Theory of Lagged and Nonlagged Responses in the Lateral Geniculate Nucleus, 1995, pp. 159–178.</ref> This spatial–temporal decorrelation makes for much more efficient coding. However, there is almost certainly much more going on.

Like other areas of the [[thalamus]], particularly other ''relay nuclei'', the LGN likely helps the [[visual system]] focus its attention on the most important information.  That is, if you hear a sound slightly to your left, the [[auditory system]] likely "tells" the [[visual system]], through the LGN via its surrounding peri-reticular nucleus, to direct visual attention to that part of space.<ref>{{Cite journal | last1 = McAlonan | first1 = K. | last2 = Cavanaugh | first2 = J. | last3 = Wurtz | first3 = R. H. | doi = 10.1523/JNEUROSCI.5602-05.2006 | title = Attentional Modulation of Thalamic Reticular Neurons | journal = Journal of Neuroscience | volume = 26 | issue = 16 | pages = 4444–4450 | year = 2006 | pmid = 16624964 | pmc = 6674014 | doi-access = free }}</ref> The LGN is also a station that refines certain [[receptive fields]].<ref>{{Cite journal | last1 = Tailby | first1 = C. | last2 = Cheong | first2 = S. K. | last3 = Pietersen | first3 = A. N. | last4 = Solomon | first4 = S. G. | last5 = Martin | first5 = P. R. | title = Colour and pattern selectivity of receptive fields in superior colliculus of marmoset monkeys | doi = 10.1113/jphysiol.2012.230409 | journal = The Journal of Physiology | volume = 590 | issue = 16 | pages = 4061–4077 | year = 2012 | pmid = 22687612 | pmc =3476648 }}</ref>

Axiomatically determined functional models of LGN cells have been determined by Lindeberg <ref name=Lin13BICY>{{cite journal | last1 = Lindeberg | first1 = T. | year = 2013 | title = A computational theory of visual receptive fields | journal = Biological Cybernetics | volume = 107 | issue = 6| pages = 589–635 | doi = 10.1007/s00422-013-0569-z | pmid = 24197240 | pmc = 3840297 }}</ref><ref name=Lin21Heliyon>{{cite journal | last1 = Lindeberg | first1 = T. | year = 2021 | title = Normative theory of visual receptive fields | journal = Heliyon | volume = 7 | issue = 1| pages = e05897 | doi=10.1016/j.heliyon.2021.e05897| pmid = 33521348 | pmc = 7820928 | doi-access = free }}</ref> in terms of Laplacian of Gaussian kernels over the spatial domain in combination with temporal derivatives of either non-causal or time-causal scale-space kernels over the temporal domain. It has been shown that this theory both leads to predictions about receptive fields with good qualitative agreement with the biological receptive field measurements performed by DeAngelis et al.<ref>{{cite journal | last1 = DeAngelis | first1 = G. C. | last2 = Ohzawa | first2 = I. | last3 = Freeman | first3 = R. D. | year = 1995 | title = Receptive field dynamics in the central visual pathways | journal = Trends Neurosci | volume = 18 | issue = 10| pages = 451–457 | doi=10.1016/0166-2236(95)94496-r | pmid=8545912| s2cid = 12827601 }}</ref><ref>G. C. DeAngelis and A. Anzai "A modern view of the classical receptive field: linear and non-linear spatio-temporal processing by V1 neurons. In: Chalupa, L.M., Werner, J.S. (eds.) The Visual Neurosciences, vol. 1, pp. 704–719. MIT Press, Cambridge, 2004.</ref> and guarantees good theoretical properties of the mathematical receptive field model, including covariance and invariance properties under natural image transformations.<ref name=Lin13PONE>{{cite journal | last1 = Lindeberg | first1 = T. | year = 2013 | title = Invariance of visual operations at the level of receptive fields | journal = PLOS ONE | volume = 8 | issue = 7| page = e66990 | doi = 10.1371/journal.pone.0066990 | pmid = 23894283 | pmc = 3716821 | arxiv = 1210.0754 | bibcode = 2013PLoSO...866990L | doi-access = free }}</ref><ref name=Lin23Front>[https://dx.doi.org/10.3389/fncom.2023.1189949 T. Lindeberg "Covariance properties under natural image transformations for the generalized Gaussian derivative model for visual receptive fields", Frontiers in Computational Neuroscience, 17:1189949, 2023.]</ref> Specifically according to this theory, non-lagged LGN cells correspond to first-order temporal derivatives, whereas lagged LGN cells correspond to second-order temporal derivatives.

===Color processing===
{{see also|Opponent process}}
The LGN is integral in the early steps of color processing, where opponent channels are created that compare signals between the different [[Photoreceptor cell]] types. The output of P-cells comprises red-green opponent signals. The output of M-cells does not include much color opponency, rather a sum of the red-green signal that evokes [[luminance]]. The output of K-cells comprises mostly blue-yellow opponent signals.<ref name=Gho17>[https://doi.org/10.1016/j.pneurobio.2017.06.002 M. Ghodrati, S.-M. Khaligh-Razavi, S.R. Lehky, Towards building a more complex view of the lateral geniculate nucleus: recent advances in understanding its role, Prog. Neurobiol. 156:214–255, 2017.]</ref><!-- This review paper can be used to update [[#Function in visual perception]] section -->

==Rodents==

In rodents, the lateral geniculate nucleus contains the dorsal lateral geniculate nucleus (dLGN), the ventral lateral geniculate nucleus (vLGN), and the region in between called the intergeniculate leaflet (IGL). These are distinct subcortical nuclei with differences in function.

===dLGN===

The dorsolateral geniculate nucleus is the main division of the lateral geniculate body. In the mouse, the area of the dLGN is about 0.48mm<math>{}^2</math>. The majority of input to the dLGN comes from the retina. It is laminated and shows retinotopic organization.<ref>{{cite journal|last=Grubb|first=Matthew S. |author2=Francesco M. Rossi |author3=Jean-Pierre Changeux |author4=Ian D. Thompson|title=Abnormal functional organization in the dorsal lateral geniculate nucleus of mice lacking the beta2 subunit of the nicotinic acetylcholine receptor|journal=Neuron|date=Dec 18, 2003|volume=40|issue=6|pages=1161–1172|doi=10.1016/s0896-6273(03)00789-x|pmid=14687550|doi-access=free}}</ref>

===vLGN===

The ventrolateral geniculate nucleus has been found to be relatively large in several species such as lizards, rodents, cows, cats, and primates.<ref>{{cite journal|last=Cooper|first=H.M.|author2=M. Herbin |author3=E. Nevo |title=Visual system of a naturally microphthalamic mammal: The blind mole rat, Spalax ehrenbergl|journal=Journal of Comparative Neurology|date=Oct 9, 2004|volume=328|issue=3|pages=313–350|doi=10.1002/cne.903280302|pmid=8440785|s2cid=28607983}}</ref>  An initial cytoarchitectural scheme, which has been confirmed in several studies, suggests that the vLGN is divided into two parts. The external and internal divisions are separated by a group of fine fibers and a zone of thinly dispersed neurons. Additionally, several studies have suggested further subdivisions of the vLGN in other species.<ref name="Harrington 1997 705–727">{{cite journal|last=Harrington|first=Mary|title=The ventral lateral geniculate nucleus and the intergeniculate leaflet: interrelated structures in the visual and circadian systems|journal=Neuroscience and Biobehavioral Reviews|date=1997|volume=21|issue=5|pages=705–727|doi=10.1016/s0149-7634(96)00019-x|pmid=9353800|s2cid=20139828}}</ref> For example, studies indicate that the cytoarchitecture of the vLGN in the cat differs from rodents. Although five subdivisions of the vLGN in the cat have been identified by some,<ref>{{cite journal|last=Jordan|first=J.|author2=H. Hollander |title=The structure of the ventral part of the lateral geniculate nucleus – a cyto- and myeloarchitectonic study in the cat|journal= Journal of Comparative Neurology|date=1972|volume=145|issue=3|pages=259–272|doi=10.1002/cne.901450302|pmid=5030906|s2cid=30586321}}</ref>  the scheme that divides the vLGN into three regions (medial, intermediate, and lateral) has been more widely accepted.

===IGL===

The intergeniculate leaflet is a relatively small area found dorsal to the vLGN. Earlier studies had referred to the IGL as the internal dorsal division of the vLGN. Several studies have described homologous regions in several species, including humans.<ref>{{cite journal|last=Moore|first=Robert Y.|title=The geniculohypothalamic tract in monkey and man|journal=Brain Research|date=1989|volume=486|pages=190–194|doi=10.1016/0006-8993(89)91294-8|pmid=2720429|issue=1|s2cid=33543381}}</ref>

The vLGN and IGL appear to be closely related based on similarities in neurochemicals, inputs and outputs, and physiological properties.

The vLGN and IGL have been reported to share many neurochemicals that are found concentrated in the cells, including neuropeptide Y, GABA, encephalin, and nitric oxide synthase. The neurochemicals serotonin, acetylcholine, histamine, dopamine, and noradrenaline have been found in the fibers of these nuclei.

Both the vLGN and IGL receive input from the retina, locus coreuleus, and raphe. Other connections that have been found to be reciprocal include the superior colliculus, pretectum, and hypothalamus, as well as other thalamic nuclei.

Physiological and behavioral studies have shown spectral-sensitive and motion-sensitive responses that vary with species. The vLGN and IGL seem to play an important role in mediating phases of the circadian rhythms that are not involved with light, as well as phase shifts that are light-dependent.<ref name="Harrington 1997 705–727"/>

==Additional images==
<gallery>
 File:Constudthal.gif|Thalamus
 File:Gray683.png|Dissection of brain-stem. Lateral view.
 File:Gray722.svg|Scheme showing central connections of the [[optic nerve]]s and optic tracts.
 File:ThalamicNuclei.svg|Thalamic nuclei
 File:ERP_-_optic_cabling.jpg|3D schematic representation of optic tracts
File:Slide2BRA.JPG|Brainstem. Posterior view.
</gallery>

== See also ==
* [[Bistratified cell]]
* [[Two-streams hypothesis]]

== References ==
{{Reflist}}

== External links ==
{{Commons category|Lateral geniculate nucleus}}
<!-- Link broken *Blohm G and Schreiber C. [http://www.auto.ucl.ac.be/EYELAB/neurophysio/light_perception/High_order_visual_proc.html LGN in the visual pathway]. Retrieved September 1, 2004. -->
* Malpeli J. [https://web.archive.org/web/20040823060304/http://soma.npa.uiuc.edu/labs/malpeli/home.html Malpeli Lab Home Page]. Retrieved September 1, 2004.
* {{BrainMaps|lateral%20geniculate%20nucleus}}
* {{UMichAtlas|eye_38}} – "The Visual Pathway from Below"
* {{BrainMaps|lgn}}
* {{MedicalMnemonics|307|640|}}

{{Visual_system}}
{{Diencephalon}}
{{Cranial nerves}}
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{{DEFAULTSORT:Lateral Geniculate Nucleus}}
[[Category:Thalamic nuclei]]
[[Category:Visual system]]