Editing Lesser kudu

{{Short description|Species of antelope}}
{{good article}}
{{speciesbox
| name = Lesser kudu
| image = Lesser Kudu Male (Tragelaphus imberbis).jpg
| image_caption = Adult male
| status = NT 
| status_system = IUCN3.1
| status_ref = <ref name=iucn>{{cite iucn|author=IUCN SSC Antelope Specialist Group|year=2016|errata=2017|title=''Tragelaphus imberbis''|page=e.T22053A115165887|doi=10.2305/IUCN.UK.2016-3.RLTS.T22053A50196563.en|access-date=12 April 2022}}</ref>
| taxon = Tragelaphus imberbis
| authority = ([[Edward Blyth|Blyth]], 1869)
| synonyms_ref = <ref name=heller/><ref name=MSW3/>
| synonyms = * ''Ammelaphus strepsiceros'' <small>(Heller, 1912)</small>
* ''Ammelaphus australis'' <small>(Heller, 1913)</small>
| range_map = Tragelaphus imberbis map.png
| range_map_caption = Range map 
}}

The '''lesser kudu''' ('''''Tragelaphus imberbis''''') is a medium-sized [[bushland]] [[antelope]] found in [[East Africa]]. The species is a part of the [[ungulate]] genus ''[[Tragelaphus]]'' (family [[Bovidae]]), along with several other related species of striped, spiral-horned African bovids, including the related [[greater kudu]], the [[Bongo (antelope)|bongo]], [[Cape bushbuck|bushbuck]], [[common eland|common]] and [[Giant eland|giant elands]], [[nyala]] and [[sitatunga]]. It was first [[Scientific description|scientifically described]] by English [[zoologist]] [[Edward Blyth]] (1869). 

The lesser kudu’s nose-to-tail length is typically {{cvt|110|-|140|cm|in}}. Males reach about {{cvt|95|-|105|cm|in}} at the shoulder, while females reach {{cvt|90|-|100|cm|in}}. Males typically weigh {{cvt|92|-|108|kg|lb}} and females {{cvt|56|-|70|kg|lb}}. [[Horn (anatomy)|Horn]]s are present only on males. The spiral horns are {{cvt|50|-|70|cm|in}} long, and have 2-2.5 complete twists. The lesser kudus have very distinctive physical markings; females and juveniles have a golden-brown coat, with white vertical stripes on their sides, while the males develop into a dark grey colour—after about two years—and grow a pronounced “streak” of shaggy hair down the centre of their backs. Males retain the golden-brown color only on the shins, with the top of each thigh having a darker black band separating it from the grey body. The inner thighs are white. Males also have distinct black “masks” on their faces, with a black underside going up to the [[sternum]]. Males and females both display a series of unique white markings and patches, including white lips, two small spots on both cheeks, a spot at the base of each ear, two spots between the eyes, white eyelids, a white throat patch, and a patch of white atop the chest. These markings may aid in camouflaging and hiding amidst shrubbery, as well as helping to cool specific areas of the body by being white.

A pure browser, the lesser kudu subsists off of foliage from tall bushes, trees (fresh branch shoots, twigs) and herbaceous perennial plants. Despite seasonal and local fluctuations, foliage from trees and shrubs constitutes 60–80% of their diet throughout the year. The lesser kudu is mainly [[crepuscular]], preferring to be active after dusk until the dawn hours, seeking shelter in dense thickets just after the sunrise. The lesser kudu exhibits no territorial behaviour, and fights are rare. While females are gregarious, adult males prefer being solitary. No fixed breeding season is seen; births may occur at any time of the year. The lesser kudu inhabits dry bushland regions, bordering on arid grasslands, as well as scrubland and light open forest habitat.

The lesser kudu is native to [[Ethiopia]], [[Kenya]], [[Somalia]], [[South Sudan]], [[Tanzania]], and [[Uganda]], but it is possibly [[Local extinction|extirpated]] from [[Djibouti]]. It may have been present in [[Saudi Arabia]] and [[Yemen]] as recently as 1967, though its presence in the Arabian Peninsula is still controversial.<ref name="kingdon" /> The total population of the lesser kudu has been estimated to be nearly 118,000, with a decreasing trend in populations. One-third of the populations survive in protected areas. Presently, the [[International Union for Conservation of Nature]]  rates the lesser kudu as "[[near threatened]]".

==Taxonomy and genetics==
{{cladogram|align=left|title=
|caption=Phylogenetic relationships of the mountain nyala from combined analysis of all molecular data (Willows-Munro et.al. 2005)
|cladogram={{clade | style=font-size:90%;line-height:100%;width:300px;
|1={{clade
|1={{clade |label1=
 |1= {{clade|label1=
           |1=
         {{clade
         |label1=
         |1={{clade
            |1={{clade
                |1={{clade
                   |1=[[Giant eland]]
                   |2=[[Common eland]]}}
                |2={{clade
                   |1=[[Greater kudu]]
                   |2={{clade
                       |1=[[Balbok]]
                       |2={{Clade
                           |1={{clade
                              |1={{clade
                                  |1=[[Bongo (antelope)|Bongo]]
                                  |2=[[Sitatunga]]}}
                           |2={{Clade
                           |1={{clade
                              |1={{clade
                                  |1=[[Cape bushbuck]]
|2=[[Harnessed bushbuck]] }} }} }} }} }} }} }} }} 
         |2=[[Nyala]]}}
|2='''Lesser kudu''' }} }} }} }} }} }}

The [[scientific name]] of the lesser kudu is ''Tragelaphus imberbis''. The animal is classified under the genus ''Tragelaphus'' in family Bovidae. It was [[Scientific description|first described]] by the English zoologist [[Edward Blyth]] in 1869.<ref name=MSW3>{{MSW3|id=14200730|page=698}}</ref> The generic name, ''Tragelaphus'', derives from [[Greek language|Greek]] word ''tragos'', meaning a male goat, and ''elaphos'', which means a deer, while the specific name ''imberbis'' comes from the Latin term meaning unbearded, referring to this kudu's lack of mane.<ref>{{MerriamWebsterDictionary|Tragelaphus|access-date=30 January 2016}}</ref> The vernacular name kudu (or koodoo) may have originated from the [[Khoikhoi language|Khoikhoi]] ''kudu'', or via the [[Afrikaans Language|Afrikaans]] ''koedoe''.<ref>{{MerriamWebsterDictionary|Kudu|access-date=30 January 2016}}</ref> The term "lesser" denotes the smaller size of this antelope as compared to the [[greater kudu]].<ref name=kingdon/>

In 1912, the genus ''Ammelaphus'' was established for just the lesser kudu by American zoologist [[Edmund Heller]], the type species being ''A. strepsiceros''.<ref name=heller>{{cite book|first=E.|last=Heller|title=New Genera and Races of African Ungulates|publisher=Smithsonian Institution|location=Washington D. C.|url=https://www.biodiversitylibrary.org/pdf3/005508900035814.pdf|page=15|date=November 2, 1912}}</ref> The lesser kudu is now typically placed in ''Tragelaphus''.<ref name=MSW3/> However, a 2011 publication by [[Colin Groves]] and [[Peter Grubb (zoologist)|Peter Grubb]] argues for the lesser kudu's renewed placement in the genus ''Ammelaphus'' on the grounds that this species is part of the earliest-diverging lineage of its tribe, having split from the main lineage before it separated into ''Tragelaphus'' and ''Taurotragus''.<ref name="Groves2011">{{cite book |last1=Groves |first1=C. |author1-link=Colin Groves |last2=Grubb |first2=P. |author2-link=Peter Grubb (zoologist) |title=Ungulate Taxonomy |date=2011 |publisher=Johns Hopkins University Press |location=Baltimore, US |isbn=978-1-4214-0093-8 |pages=139}}</ref>

In 2005, Sandi Willows-Munro (of the [[University of KwaZulu-Natal]]) and colleagues carried out a [[mitochondria]]l analysis of the nine ''Tragelaphus'' species. [[mtDNA]] and [[nDNA]] data were compared. The results showed that the tribe Tragelaphini is [[monophyletic]] with the lesser kudu [[Basal (phylogenetics)|basal]] in the phylogeny, followed by the [[nyala]] (''T. angasii'').<ref name=cladogram>{{cite journal|last1=Willows-Munro|first1=S.|last2=Robinson|first2=T. J.|last3=Matthee|first3=C. A.|title=Utility of nuclear DNA intron markers at lower taxonomic levels: Phylogenetic resolution among nine ''Tragelaphus'' spp.|journal=Molecular Phylogenetics and Evolution|date=June 2005|volume=35|issue=3|pages=624–36|doi=10.1016/j.ympev.2005.01.018|pmid=15878131|bibcode=2005MolPE..35..624W }}</ref><ref name="groves2014">{{cite journal|last1=Groves|first1=C.|title=Current taxonomy and diversity of crown ruminants above the species level|journal=Zitteliana|date=2014|volume=32|issue=B|pages=5–14|url=http://www.palmuc.de/bspg/images/pdf/zitteliana32/1_groves.pdf|issn=1612-4138|access-date=2016-01-30|archive-url=https://web.archive.org/web/20160204110133/http://www.palmuc.de/bspg/images/pdf/zitteliana32/1_groves.pdf|archive-date=2016-02-04|url-status=dead}}</ref> On the basis of mitochondrial data,  the lesser kudu separated from its [[sister clade]] around 13.7 million years ago. However, the nuclear data show that lesser kudu and nyala form a clade, and collectively separated from the sister clade 13.8 million years ago.<ref name="ropiquet">{{cite journal|last1=Ropiquet|first1=A.|title=Etude des radiations adaptatives au sein des Antilopinae (Mammalia, Bovidae)|journal=Ph.D. Thesis, Université Paris|date=2006|volume=6|issue=1–247}}</ref><ref name="hassanin">{{cite journal|last1=Hassanin|first1=A.|last2=Delsuc|first2=F.|last3=Ropiquet|first3=A.|last4=Hammer|first4=C.|last5=Jansen van Vuuren|first5=B.|last6=Matthee|first6=C.|last7=Ruiz-Garcia|first7=M.|last8=Catzeflis|first8=F.|last9=Areskoug|first9=V.|last10=Nguyen|first10=T.T.|last11=Couloux|first11=A.|title=Pattern and timing of diversification of Cetartiodactyla (Mammalia, Laurasiatheria), as revealed by a comprehensive analysis of mitochondrial genomes|journal=Comptes Rendus Biologies|date=2012|volume=335|issue=1|pages=32–50|doi=10.1016/j.crvi.2011.11.002|pmid=22226162|doi-access=free}}</ref>

The lesser kudu has 38 [[Diploidy|diploid]] [[chromosome]]s. However, unlike others in the [[Subfamily (biology)|subfamily]] Tragelaphinae, the [[X chromosome]] and [[Y chromosome]] are compound and each is fused with one of two identical [[autosome]]s.<ref>{{cite journal|last1=Benirschke|first1=K.|last2=Ruedi|first2=D.|last3=Muller|first3=H.|last4=Kumamoto|first4=A.T.|last5=Wagner|first5=K.L.|last6=Downes|first6=H.S.|title=The unusual karyotype of the lesser kudu,''Tragelaphus imberbis''|journal=Cytogenetic and Genome Research|date=1980|volume=26|issue=2–4|pages=85–92|doi=10.1159/000131429|pmid=7389415}}</ref>

==Physical description==
[[File:Male Lesser Kudu.jpg|thumbnail|left|Male lesser kudu]]
The lesser kudu is a spiral-horned antelope. The head-and-body length is typically between {{cvt|110|and|140|cm|in}}. Males reach about {{cvt|95|-|105|cm|in}} at the shoulder, while females reach {{cvt|90|-|100|cm|in}}. Males typically weigh {{cvt|92|-|108|kg|lb}} and females {{cvt|56|-|70|kg|lb}}.<ref name=estes>{{cite book|last=Estes|first=R. D.|title=The Behavior Guide to African Mammals : Including Hoofed Mammals, Carnivores, Primates|date=2004|publisher=University of California Press|location=Berkeley|isbn=0520080858|pages=[https://archive.org/details/isbn_0520080858/page/180 180–2]|edition=4th|url=https://archive.org/details/isbn_0520080858/page/180}}</ref> The bushy tail is {{cvt|25|-|40|cm|in}} long, white underneath and with a black tip at the end.<ref name=adw>{{cite web|last=Paschka|first=N.|title=''Tragelaphus imberbis'' (lesser kudu)|url=http://animaldiversity.ummz.umich.edu/accounts/Tragelaphus_imberbis/|work=University of Michigan Museum of Zoology|publisher=Animal Diversity Web|access-date=2 March 2014}}</ref>

Distinct signs of [[sexual dimorphism]] are seen in the antelope. The male is considerably larger than the female. The females, as well as juveniles, have a [[rufous]] coat, whereas the males become yellowish grey or darker after the age of 2 years. The male has a prominent black crest of hair on the neck, but this feature is not well-developed in the female.<ref name=kingdon>{{cite book|last=Kingdon|first=J.|title=Mammals of Africa|date=2013|publisher=Bloomsbury Publishing|location=London|isbn=978-1408189962|pages=142–7|author2=Butynski, T. |author3=Happold, D. }}</ref> One long white stripe runs along the back, with 11–14 white stripes branching towards the sides.<ref name=adw/> The chest has a central black stripe, and  no throat beard is present.<ref name=estes/> A black stripe runs from each eye to the nose and a white one from each eye to the centre of the dark face. A [[Chevron (anatomy)|chevron]] is present between the eyes. The area around the lips is white, the throat has white patches, and two white spots appear on each side of the lower jaw. The underparts are completely white, while the slender legs are tawny and have black and white patches.<ref name=adw/> The lesser kudu is characterised by large, rounded ears. Its tracks are similar to the greater kudu's.<ref name=track>{{cite book|last=Chris|first=S.|title=A Field Guide to the Tracks and Signs of Southern and East African Wildlife|date=2000|publisher=Struik|location=Cape Town|isbn=1868725588|edition=3rd|author2=Stuart, T.|url=https://books.google.com/books?id=WcDPj5hgT30C}}{{Dead link|date=August 2024 |bot=InternetArchiveBot |fix-attempted=yes }}</ref> Females have four teats.<ref name=kingdon/> The average lifespan is 10 years in the wild, and 15 years in captivity.<ref name=adw/>

Horns are present only on males. The spiral horns are {{cvt|50|-|70|cm|in}} long, and have two to two-and-a-half twists.<ref name=estes/> The base circumference is {{cvt|156|-|171|cm|in}}.<ref name=adw/> The slender horns are dark brown and tipped with white.<ref name=kingdon/> Male young begin developing horns after 6-8 months, which reach full length after 3 years.<ref name=adw/>

==Ecology and behaviour==
[[File:Tragelaphus imbersis Dvur zoo 4.jpg|thumbnail|Herd of lesser kudu in [[Dvůr Králové Zoo]]]]
The lesser kudu is mainly active at night and during the dawn, and seeks shelter in dense thickets just after the sunrise.<ref name=adw/> It can [[camouflage]] so well in such dense vegetation that only its ears and tail can indicate its presence.<ref name=ark>{{cite web|title=Lesser kudu|url=http://www.arkive.org/lesser-kudu/tragelaphus-imberbis/|work=Wildscreen|publisher=ARKive|access-date=2 March 2014|archive-url=https://web.archive.org/web/20140304165605/http://www.arkive.org/lesser-kudu/tragelaphus-imberbis/|archive-date=2014-03-04|url-status=dead}}</ref> The midday is spent in rest and rumination in shaded areas.<ref name=kingdon/><ref name=mitchell>{{cite journal|last=Mitchell|first=A. W.|title=Preliminary observations on the daytime activity patterns of lesser kudu in Tsavo National Park, Kenya|journal=African Journal of Ecology|date=September 1977|volume=15|issue=3|pages=199–206|doi=10.1111/j.1365-2028.1977.tb00398.x|bibcode=1977AfJEc..15..199M }}</ref> The animal spends about equal time foraging, standing and lying, and roaming.<ref name=estes/> As a thin tragelaphine, the lesser kudu can move readily through dense vegetation with ease. The lesser kudu is a shy and wary animal. When alarmed, the animal stands motionless. If it senses any approaching predator, it gives out a short sharp bark, similar to the [[Cape bushbuck|bushbuck]]'s,  then makes multiple leaps up to {{cvt|2|m|ft|abbr=on}} high with an upraised tail. If captured by the predator, the victim gives a loud bleat.<ref name=estes/>

Lesser kudus are gregarious in nature, especially females. No distinct leader or any hierarchy is noted in the social structure; with no territorial behavior, fights are uncommon. While fighting, the lesser kudus interlock horns and try pushing one another. Mutual grooming is hardly observed.<ref name=kingdon/> Unlike most tragelaphines, females can be closely associated for several years. One to three females, along with their offspring, may form a group. Juvenile males leave their mothers when aged a year and a half, and may form pairs. However, at the age of 4-5 years, males prefer a solitary lifestyle and avoid one another, though four or five bulls may share the same home range. Lesser kudu do not usually associate with other animals, except when they feed in the same area.<ref name=estes/><ref name=ark/>

===Diet===
[[File:Lesser Kudu Female.jpg|thumbnail|Feeding female and juvenile]]
A pure browser, the lesser kudu feeds on foliage from bushes and trees (shoots, twigs) and herbs.<ref name=adw/> It also eats flowers and fruits if available, and takes small proportions of grasses, usually in the wet season. Despite seasonal and local variations, foliage from trees and shrubs constitutes 60-80% of the diet throughout the year. Foliage from creepers and vines (such as ''Thunbergia guerkeana'' and some species of [[Cucurbitaceae]] and [[Convulvulaceae]]) forms 15-25% of the diet in the wet season. Fruits are consumed mainly in the dry season. [[Olfactory]] searching, much in the same posture as grazing, is used to find fallen fruits (such as ''Melia volkensii'' and ''[[Acacia tortilis]]''), while small fruits (such as ''[[Commiphora]]'' species) are directly plucked from trees. The size and structure of its stomach also suggests its primary dependence on browse.<ref name=kingdon/>

The lesser kudu browses primarily at dusk or dawn, or nocturnally, <ref name=mitchell/> and is sometimes associated with [[gerenuk]] and the [[impala]].<ref name=kingdon/> The lesser kudu and the gerenuk might compete for evergreen species in the dry season.<ref name=kingdon/> However, unlike the long-necked gerenuk, the lesser kudu rarely consumes ''[[Acacia]]'' species, and does not stand on its hindlegs while feeding.<ref name=estes/> The lesser kudu likewise does not have a great requirement for water, and can thrive in arid environments<ref name=adw/> as it is able to extract sufficient moisture from [[Succulent plant|succulent plants]], such as the leaves of wild [[sisal]] and ''[[Sansevieria]]'' ('snake-plants', genus ''[[Dracaena (plant)|Dracaena]]''), and certain species of the poisonous [[Euphorbiaceae]] family; it drinks water readily when rains come or when sources are available.<ref name=kingdon/><ref name=estes/>

===Reproduction===
Both the males and females become sexually mature by the time they are a year and a half old. However, males actually mate after the age of four to five years.<ref name=adw/> Males and females are most reproductive till the age of 14 and 14–18 years, respectively, with the maximum age of successful lactation in females being 13–14 years.<ref name=rep/> With no fixed breeding season, births may occur at any time of the year. A study at [[Dvůr Králové Zoo]] ([[Czech Republic]]) showed that 55% of the births occurred between September and December.<ref name=rep>{{cite journal|last=Váhala|first=J.|title=Reproduction of the lesser kudu (''Tragelaphus imberbis'') at Dvůr Králové Zoo|journal=Zoo Biology|date=1992|volume=11|issue=2|pages=99–106|doi=10.1002/zoo.1430110205}}</ref> A [[Rut (mammalian reproduction)|rutting]] male tests the urine of any female he encounters, to which the female responds by urinating. Having located a female in [[estrus]], the male follows her closely, trying to rub his cheek on her rump, head, neck, and chest. He performs gasping movements with his lips. Finally, the male mounts the female, resting his head and neck on her back, in a similar way as other tragelaphines.<ref name=kingdon/><ref name=estes/>

The gestational period is 7-8 months, after which a single calf is born. A female about to give birth isolates herself from her group, and remains alone for some days afterward. The newborn calf weighs {{convert|4|-|7.5|kg|lb|abbr=on}}. Around 50% of the calves die within the first six months of birth, and only 25% can survive after three years. In a study at [[Basle Zoo]] ([[Switzerland]]), where 43% of the offspring from captive breeding died before reaching the age of six months, the major causes of high juvenile mortality were found to be the spread of [[white muscle disease]] and deficiency of [[vitamin E]] and [[selenium]] in diets. The herd size, sex, interbreeding, and season did not play any role in juvenile mortality.<ref>{{cite journal|last=Besselmann|first=D.|author2=Schaub, D.|author3=Wenker, C.|author4=Völlm, J.|author5=Robert, N.|author6=Schelling, C.|author7=Steinmetz, H.|author8=Clauss, M.|title=Juvenile mortality in captive lesser kudu (''Tragelaphus imberbis'') at Basle Zoo and its relation to nutrition and husbandry|journal=Journal of Zoo and Wildlife Medicine|date=March 2008|volume=39|issue=1|pages=86–91|doi=10.1638/2007-0004.1|pmid=18432100|s2cid=22836125|url=https://www.zora.uzh.ch/id/eprint/3424/5/JZWM_Besselmann_kudu_2008V.pdf|access-date=2019-12-12|archive-date=2017-09-24|archive-url=https://web.archive.org/web/20170924143612/http://www.zora.uzh.ch/id/eprint/3424/5/JZWM_Besselmann_kudu_2008V.pdf|url-status=dead}}</ref> The mother hides her calf while she goes out to feed, and returns mainly in the evening to suckle her young. She checks the calf's identity by sniffing its rump or neck. In the first month, suckling may occur for 8 minutes. The mother and calf communicate with low bleats. She licks her offspring, particularly in the [[Perineal body|perineal region]], and may consume its excreta.<ref name=kingdon/><ref name=estes/>

==Habitat and distribution==
The lesser kudu inhabits dry bushland regions.<ref name=adw/> It is closely associated with ''Acacia'' and ''Commiphora'' thornbush in semiarid areas of northeastern Africa. The animal avoids open areas and long grass, preferring shaded areas with short grasses instead.<ref name=iucn/> Found in woodlands and hilly areas, as well, the lesser kudu is generally found at altitudes below {{cvt|1200|m|ft|abbr=on}};<ref name=estes/> though they have been recorded at heights about {{cvt|1740|m|ft|abbr=on}} near [[Mount Kilimanjaro]].<ref name=iucn/> While individual home ranges of these animals are {{cvt|0.4|-|6.7|km2|sqft|lk=in|sigfig=2|abbr=on}} in size, those of males have an average size of {{convert|2.2|km2|sqft|lk=in|sigfig=2|abbr=on}} and those of females {{convert|1.8|km2|sqft|lk=in|sigfig=2|abbr=on}}.<ref name=nowak/>

The lesser kudu is native to [[Ethiopia]], [[Kenya]], [[Somalia]], [[South Sudan]], [[Tanzania]], and [[Uganda]], but it is extinct in [[Djibouti]].<ref name=iucn/> Largely confined to the [[Horn of Africa]] today, the species historically ranged from [[Awash, Ethiopia|Awash]] (Ethiopia) southward through southern and eastern Ethiopia, and most parts of Somalia (except the north and the northeast) and Kenya (except the southwest). It also occurred in southeastern Sudan and northeastern and eastern parts of Uganda and Tanzania. Evidence for its existence in the [[Arabian Peninsula]] includes a set of horns obtained in 1967 from an individual shot in [[South Yemen]] and another in [[Saudi Arabia]], as well as a recent analysis of  early and middle [[Holocene]] [[rock art]] sites in [[Rock Art in the Ha'il Region|Jubbah and Shuwaymis]], [[Ha'il province]], Saudi Arabia.<ref name=kingdon/><ref>Guagnin M, Shipton C, el-Dossary S, et al. Rock art provides new evidence on the biogeography of kudu (Tragelaphus imberbis), wild dromedary, aurochs (Bos primigenius) and African wild ass (Equus africanus) in the early and middle Holocene of north-western Arabia. J Biogeogr. 2018;00:1–14. https://doi.org/10.1111/jbi.13165</ref>

==Threats and conservation==
The lesser kudu's shyness and its ability to camouflage itself in dense cover has protected it from the risks of poaching. For instance, the lesser kudu is widespread in the Ogaden region, which is rich in dense bush, despite reckless hunting by local people.<ref name=iucn/> However, [[rinderpest]] outbreaks, to which the lesser kudu is highly susceptible, have resulted in a steep decline of 60% in the animal's population in Tsavo National Park in Kenya.<ref>{{cite book|last=Sherman|first=D. M.|title=Tending Animals in the Global Village: A Guide to International Veterinary Medicine|date=2002|publisher=Lippincott Williams & Wilkins|location=Philadelphia|isbn=0470292105|page=264}}</ref> [[Overgrazing]], human settlement, and loss of habitat are some other threats to the survival of the lesser kudu.<ref name=iucn/>

The total population of the lesser kudu has been estimated to be nearly 118,000, with a decreasing trend in populations. The rate of decline has increased to 20% over two decades. Presently, the [[IUCN]] rates the lesser kudu as "[[near threatened]]".<ref name=iucn/> Around a third of the population of the lesser kudu occurs in protected areas such as [[Awash, Ethiopia|Awash]], [[Omo National Park|Omo]] and [[Mago National Park|Mago]] National Parks (Ethiopia); [[Lag Badana National Park]] (Somalia); [[Tsavo|Tsavo National Park]] (Kenya); [[Ruaha National Park]] and game reserves (Tanzania), though it occurs in larger numbers outside these areas.<ref name=east>{{cite book|last1=East|first1=R.|last2=IUCN/SSC |first2=Antelope Specialist Group|title=African Antelope Database 1998|date=1999|publisher=The IUCN Species Survival Commission|location=Gland, Switzerland|isbn=2831704774|pages=132–4}}</ref> Population density rarely exceeds 1/km<sup>2</sup>., and is generally much lower.<ref name=nowak>{{cite book|last=Nowak|first=R. M.|title=Walker's Mammals of the World|url=https://archive.org/details/walkersmammalsof0002nowa|url-access=registration|date=1999|publisher=Johns Hopkins University Press|location=Baltimore, Maryland|isbn=0801857899|pages=[https://archive.org/details/walkersmammalsof0002nowa/page/1140 1140–1]|edition=6th}}</ref>

The handsome head of the male lesser kudu, with his elegant spiraled horns, is the symbol of the [[Saint Louis Zoo]].

==References==
{{Reflist|30em}}

==External links==
{{portal|Mammals}}
* Information from [http://www.awf.org/wildlife-conservation/kudu African Wildlife Foundation]
*{{Commonscat-inline|Tragelaphus imberbis}}
*{{Wikispecies-inline|''Tragelaphus imberbis''}}

{{Artiodactyla|R.4}}
{{Taxonbar|from=Q622882}}

[[Category:Tragelaphus|lesser kudu]]
[[Category:Fauna of East Africa]]
[[Category:Fauna of the Horn of Africa]]
[[Category:Mammals of Ethiopia]]
[[Category:Mammals of Kenya]]
[[Category:Mammals of Somalia]]
[[Category:Mammals of South Sudan]]
[[Category:Mammals of Tanzania]]
[[Category:Mammals of Uganda]]
[[Category:Mammals described in 1869|lesser kudu]]
[[Category:Taxa named by Edward Blyth|lesser kudu]]
[[Category:Bovids of Africa]]