Editing Manis

{{Short description|Genus of mammals}}
{{Other uses}}
{{Automatic taxobox
| name = Asiatic pangolin
| fossil_range = {{fossil_range|12.9|0}} Middle [[Miocene]] - present<ref name="Gaubert2018">Philippe Gaubert, Agostinho Antunes, Hao Meng, Lin Miao, Stéphane Peigné, Fabienne Justy, Flobert Njiokou, Sylvain Dufour, Emmanuel Danquah, Jayanthi Alahakoon, Erik Verheyen, William T Stanley, Stephen J O’Brien, Warren E Johnson, Shu-Jin Luo (2018) [https://academic.oup.com/jhered/article/109/4/347/4622594 "The Complete Phylogeny of Pangolins: Scaling Up Resources for the Molecular Tracing of the Most Trafficked Mammals on Earth"] Journal of Heredity, Volume 109, Issue 4, Pages 347–359</ref>
| image = Manis.jpg
| image_caption = Living pangolins from genus ''Manis''
| taxon = Manis
| parent_authority = Gray, 1821
| authority = [[Carl Linnaeus|Linnaeus]], [[10th edition of Systema Naturae|1758]]
| type_species = ''[[Chinese pangolin|Manis pentadactyla]]''
| type_species_authority = Linnaeus, 1758
| subdivision_ranks = Species
| subdivision =
* ''[[Manis aurita]]''
* ''[[Indian pangolin|Manis crassicaudata]]''
* ''[[Chinese pangolin|Manis pentadactyla]]''
* †''[[Manis hungarica]]''
* †''[[Manis lydekkeri]]''
* Subgenus: '''''Paramanis'''''
** ''[[Philippine pangolin|Manis culionensis]]''
** ''[[Sunda pangolin|Manis javanica]]''
** ''[[Manis mysteria]]''
** †''[[Manis palaeojavanica]]''
| range_map = Manis ranges.png
| range_map_caption = {{leftlegend|#7C1797|''[[Indian pangolin|Manis crassicaudata]]''|outline=black}}
{{leftlegend|#FF0800|''[[Philippine pangolin|Manis culionensis]]''|outline=black}}
{{leftlegend|#59C9FD|''[[Sunda pangolin|Manis javanica]]''|outline=black}}
{{leftlegend|#F89B3C|''[[Chinese pangolin|Manis pentadactyla]]''|outline=black}}
| synonyms = {{collapsible list |bullets=true |title=synonyms of subfamily:
|Mania
|Manidae {{small|(Gray, 1821)}}
|Manina {{small|(Gray, 1825)}}<ref>J. E. Gray. (1825.) "An outline of an attempt at the disposition of Mammalia into Tribes and Families, with a list of genera apparently appertaining to each Tribe." Annals of Philosophy, new series 10:337-344</ref>
|Pholidotina {{small|(Gray, 1873)}}
}}
{{collapsible list |bullets=true |title=synonyms of genus:
|''Pangolin'' {{small|(Gray, 1873)}}
|''Pangolinus'' {{small|(Rafinesque, 1821)}}
|''Paramanis'' {{small|(Pocock, 1924)}}
|''Phatages'' {{small|(Sundevall, 1843)}}
|''Phatagenus'' {{small|(Sundevall, 1843)}}
|''Pholidotus'' {{small|(Brisson, 1762)}}
|''Quaggelo'' {{small|(Frisch, 1775)}}
}}
| synonyms_ref = <ref>{{Cite book |url=https://archive.org/details/indexgenerummam00palmgoog |page=[https://archive.org/details/indexgenerummam00palmgoog/page/n827 822] |title=Index Generum Mammalium: A List of the Genera and Families of Mammals |last=Palmer |first=Theodore Sherman |date=1904 |publisher=U. S. Government Printing Office |language=en}}</ref><ref>{{Cite web |url=https://www.mv.helsinki.fi/home/mhaaramo/metazoa/deuterostoma/chordata/synapsida/eutheria/leptictida/history_Manis.html |title=Taxonomic history of the genus ''Manis'' |website=Mikko's Phylogeny Archive |access-date=2019-09-18}}</ref>
}}

'''''Manis'''''  ("[[Manes|spirit]]") is a [[genus]] of [[South Asia]]n and [[East Asia]]n [[pangolin]]s, the '''Asiatic pangolins''', from subfamily Maninae, within family [[Manidae]].<ref>{{Catalogue of Life|d36852dde74a91d7c965591b175947ec}}</ref><ref name="Schlitter">{{MSW3 Pholidota |id=13900003 |pages=530-531}}</ref>

==Taxonomy==
* Subfamily: '''Maninae''' (Asiatic pangolins)
** Genus: '''''Manis''''' (Asiatic pangolins)
*** ''[[Indian pangolin|Manis crassicaudata]]'' (Indian pangolin)
*** ''[[Chinese pangolin|Manis pentadactyla]]'' (Chinese pangolin)
*** ''[[Manis aurita]]''<ref name="Zijlstra-2025">{{cite journal|vauthors=Zijlstra JS|title=''Manis aurita'' Hodgson 1837 as a valid species of pangolin: a comment to Wangmo et al. (2025)|journal=Mammalian Biology|date=25 April 2025|doi=10.1007/s42991-025-00495-x}}</ref> (= ''Manis indoburmanica'')<ref name = "Wangmo et al., 2025">Wangmo, L. K., Ghosh, A., Dolker, S., Joshi, B. D., Sharma, L. K., & Thakur, M. (2025). [https://link.springer.com/article/10.1007/s42991-024-00475-7 Indo-Burmese pangolin (''Manis indoburmanica''): a novel phylogenetic species of pangolin evolved in Asia.] Mammalian Biology, 1-8.</ref>
*** ''[[Manis mysteria]]'' (Cryptic pangolin)<ref>https://www.sci.news/biology/manis-mysteria-12316.html</ref>
*** †''[[Manis hungarica]]''
*** †''[[Manis lydekkeri]]''
*** Subgenus: '''''Paramanis'''''
**** ''[[Philippine pangolin|Manis culionensis]]'' (Philippine pangolin)
**** ''[[Sunda pangolin|Manis javanica]]'' (Sunda pangolin)
**** †''[[Manis palaeojavanica]]'' (Giant asian pangolin)

==Phylogeny==
Phylogenetic position of genus ''Manis'' within family Manidae.<ref name="web2.utc.edu">{{cite journal |last=Gaudin |first=Timothy |year=2009 |title=The Phylogeny of Living and Extinct Pangolins (Mammalia, Pholidota) and Associated Taxa: A Morphology Based Analysis |journal=Journal of Mammalian Evolution |publisher=Springer Science+Business Media |location=Heidelberg, Germany |volume=16 |issue=4 |pages=235–305 |doi=10.1007/s10914-009-9119-9 |s2cid=1773698 |url=http://web2.utc.edu/~gvv824/Gaudin%20et%20al%202009.pdf |access-date=2020-11-19 |archive-date=2015-09-25 |archive-url=https://web.archive.org/web/20150925134257/http://web2.utc.edu/~gvv824/Gaudin%20et%20al%202009.pdf |url-status=dead }}</ref><ref name="KondrashovAgadjanian2012">{{cite journal |last1=Kondrashov |first1=Peter |last2=Agadjanian |first2=Alexandre K. |title=A nearly complete skeleton of ''Ernanodon'' (Mammalia, Palaeanodonta) from Mongolia: morphofunctional analysis |journal=Journal of Vertebrate Paleontology |volume=32 |issue=5 |year=2012 |issn=0272-4634 |doi=10.1080/02724634.2012.694319 |pages=983–1001|bibcode=2012JVPal..32..983K |s2cid=86059673 }}</ref><ref name="Gaubert2018"/>
{{clade |style=font-size:85%;line-height:100%;
|label1='''[[Pholidotamorpha]]'''|sublabel1={{small|('''Pholidota''' ''sensu lato'')}}
|1={{clade
   |1=†'''[[Palaeanodonta]]''' [[File:Metacheiromys_DB152-2.jpg|60px]]
   |label2='''[[Pangolin|Pholidota]]'''|sublabel2=''sensu stricto''
   |2={{clade
      |thickness2=2
      |1=†''[[Euromanis]]''
      |label2='''?'''|state2=dotted
      |2=†Pholidota sp. (''BC 16’08'')
      |3={{clade
         |1=†'''[[Eurotamandua|Eurotamanduidae]]'''
         |label2='''[[Eupholidota]]'''
         |2={{clade
            |1=†'''[[Eomanis|Eomanoidea]]'''
            |label2='''[[Manoidea]]'''
            |2={{clade
               |thickness2=2
               |1=†'''[[Patriomanidae]]'''
               |label2='''?'''|state2=dotted
               |2=†''[[Necromanis]]''
               |label3='''[[Manidae]]'''
               |3={{clade
                  |thickness2=2 |thickness3=2 |label1='''Smutsiinae'''|sublabel1=''sensu lato''
                  |1={{clade
                     |1='''[[Phataginus|Phatagininae]]''' [[File:Anatomische Untersuchungen über die Edentaten (1852) Phataginus tricuspis.png|90px]]
                     |2=[[Smutsia|'''Smutsiinae''' {{small|(''sensu stricto'')}}]] [[File:Cambridge Natural History Mammalia Fig 109.jpg|60px]]
                     }}
                  |label2='''?'''|state2=dotted
                  |2=†[[Fayum pangolin|Manidae sp. (''DPC 3972'' & ''DPC 4364'')]]
                  |label3='''Maninae'''
                  |3={{clade
                     |thickness=2 |label1='''''Manis'''''
                     |1={{clade
                        |thickness=2
                        |1={{clade
                           |thickness=2
                           |1=''[[Chinese pangolin|Manis pentadactyla]]'' [[File:Pangolin Hardwicke (white background).jpg|60px]]
                           |label2='''?'''|state2=dotted
                           |2=†''[[Manis hungarica]]''
                           }}
                        |2={{clade
                           |thickness=2
                           |1={{clade
                              |thickness=2
                              |1=''[[Indian pangolin|Manis crassicaudata]]''
                              |label2='''?'''|state2=dotted
                              |2=†''[[Manis lydekkeri]]''
                              }}
                           |2={{clade
                              |thickness=2 |label1='''?'''|state1=dotted
                              |1=''Manis'' sp. (''Scale_H4'' & ''Scale_H8'')
                              |label2=('''''Para[[manis]]''''')
                              |2={{clade
                                 |thickness=2
                                 |1=''[[Philippine pangolin|Manis culionensis]]''
                                 |2=''[[Sunda pangolin|Manis javanica]]'' [[File:Anatomische Untersuchungen über die Edentaten (1852) Manis javanica.jpg|80px]]
                                 |label3='''?'''|state3=dotted
                                 |3=†''[[Manis palaeojavanica]]''
                                 }} }} }} }} }} }} }} }} }} }} }} }}

==References==
{{Reflist}}

{{Pholidota}}
{{Pholidotamorpha|state=autocollapse}}
{{Taxonbar|from=Q25397}}
{{Authority control}}

[[Category:Manis| ]]
[[Category:Mammal genera]]
[[Category:Taxa named by Carl Linnaeus]]
[[Category:Mammals of Asia]]


{{mammal-stub}}