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Old World flycatcher
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{{Short description|Family of birds}} {{About|the old world flycatcher family ''Muscicapidae''|the new world Tyrant Flycatcher family ''Tyrannidae''|Tyrant flycatcher||Flycatcher (disambiguation){{!}}Flycatcher}} {{Automatic taxobox | name = Old World flycatchers | image = White-eyed slaty flycatcher.jpg | image_upright = 1.2 | image_caption = [[White-eyed slaty flycatcher]],<br />(''Melaenornis fischeri'') | taxon = Muscicapidae | authority = [[John Fleming (naturalist)|Fleming J.]], 1822 | subdivision_ranks = Genera | subdivision = See text }} The '''Old World flycatchers''' are a large family, the '''Muscicapidae''', of small [[passerine]] [[bird]]s restricted to the [[Old World]] ([[Europe]], [[Africa]] and [[Asia]]), with the exception of several vagrants and two species, [[bluethroat]] (''Luscinia svecica'') and [[northern wheatear]] (''Oenanthe oenanthe''), found also in North America. These are mainly small arboreal [[insectivore]]s, many of which, as the name implies, take their [[prey]] on the wing. The family is relatively large and includes 357 species, which are divided into 57 [[genera]]. ==Taxonomy== The name Muscicapa for the family was introduced by the Scottish naturalist [[John Fleming (naturalist)|John Fleming]] in 1822.<ref>{{ cite book | last=Fleming | first=John | author-link=John Fleming (naturalist)| year=1822 | title=The philosophy of zoology; or a general view of the structure, functions, and classification of animals. | volume=2 | publisher=Hurst, Robinson & Co | place=Edinburgh | page=240 | url=https://www.biodiversitylibrary.org/page/28230398 }}</ref><ref>{{cite book | last=Bock | first=Walter J. | year=1994 | title=History and nomenclature of avian family-group names | series=Bulletin of the American Museum of Natural History Issue 222 | page=116 | hdl=2246/830 }}</ref> The word had earlier been used for the genus ''[[Muscicapa]]'' by the French zoologist [[Mathurin Jacques Brisson]] in 1760.<ref>{{ cite book | last=Brisson | first=Mathurin Jacques | author-link=Mathurin Jacques Brisson | year=1760 | title=Ornithologie; ou, Méthode contenant la division des oiseaux en ordres, sections, genres, espéces & leurs variétés. &c. | publisher=Jean-Baptiste Bauche | place=Paris | at=[https://www.biodiversitylibrary.org/page/36010429 Vol. 1 p. 32], [https://www.biodiversitylibrary.org/page/36011661 Vol. 2 p. 357] | language=la, fr}}</ref> Muscicapa comes from the Latin ''[[wikt:musca|musca]]'' meaning a fly, and ''[[wikt:capio|capere]]'' to catch.<ref>{{cite book | last= Jobling | first= James A. | year= 2010| title= The Helm Dictionary of Scientific Bird Names | url= https://archive.org/details/Helm_Dictionary_of_Scientific_Bird_Names_by_James_A._Jobling | publisher=Christopher Helm | location = London, United Kingdom | isbn = 978-1-4081-2501-4 | page=[https://archive.org/details/Helm_Dictionary_of_Scientific_Bird_Names_by_James_A._Jobling/page/n260 260]}}</ref><ref>{{L&S|musca1|musca}}, {{L&S|capio1|capere|ref}}</ref> In 1910, the German ornithologist [[Ernst Hartert]] found it impossible to define boundaries between the three families Muscicapidae, [[Sylviidae]] (Old World warblers) and [[Turdidae]] (thrushes). He therefore treated them as subfamilies of an extended flycatcher family that also included [[Timaliidae]] (Old World babblers) and [[Monarchidae]] (Monarch flycatchers).<ref>{{ cite book | last=Hartert | first=Ernst | year=1910 | title=Die Vögel der paläarktischen Fauna systematische Übersicht der in Europa, Nord-Asien und der Mittelmeerregion vorkommenden Vögel. | volume=1 | page=469 | publisher=R. Friedländer & Sohn | place=Berlin | language=de | url=https://www.biodiversitylibrary.org/page/14030276 }}</ref><ref>{{cite book| last=Taylor | first=B. | chapter=Old World Flycatchers (Muscicapidae) | editor1-last=del Hoyo | editor1-first=J. | editor2-last=Elliott | editor2-first=A. | editor3-last=Sargatal | editor3-first=J. | editor4-last=Christie | editor4-first=D.A. | editor5-last=de Juana | editor5-first=E. | title=Handbook of the Birds of the World Alive | year=2020 | publisher=Lynx Edicions | doi=10.2173/bow.muscic3.01 | s2cid=216288554 | chapter-url=http://www.hbw.com/node/52322 | access-date=30 May 2016}}{{subscription required}}</ref> Forty years later, a similar arrangement was adopted by the American ornithologists [[Ernst Mayr]] and [[Dean Amadon]] in an article published in 1951. Their large family, Muscicapidae, which they termed the "primitive insect eaters" contained 1460 species divided into eight subfamilies.<ref>{{ cite book | last1=Mayr | first1=E. | author1-link=Ernst Mayr | last2=Amadon | first2=D. | author2-link=Dean Amadon | year=1951 | title=A Classification of Recent Birds | series=American Museum Novitates, Number 1496 | publisher=American Museum of Natural History | place=New York | pages=17–19, 36–37 | hdl=2246/3994 }}</ref> The use of the extended group was endorsed by a committee set up following the Eleventh International Ornithological Congress held in Basel in 1954.<ref>{{ cite journal | last1=Mayr | first1=E. | last2=Greenway | first2=J.C. Jr. | year=1956 | title=Sequence of passerine families (Aves) | journal=Breviora | publisher=Museum of Comparative Zoology, Harvard | volume=58 | pages=1–11 | url=https://www.biodiversitylibrary.org/page/3189827 }}</ref> Subsequent [[DNA–DNA hybridization]] studies by [[Charles Sibley]] and others showed that the subfamilies were not closely related to one another. As a result, the large group was broken up into a number of separate families,<ref>{{ cite book | editor1-last=Mayr | editor1-first=E. | editor2-last=William | editor2-first=C.G. | year=1986 | title=Check-list of Birds of the World. | volume=11 | publisher=Museum of Comparative Zoology | place=Cambridge, Massachusetts | pages=v–vi | url=https://www.biodiversitylibrary.org/page/14483694 }}</ref> although for a while most authorities continued to retain the thrushes in Muscicapidae.<ref>{{ cite book | last1=Sibley | first1=C.G. | last2=Monroe | first2=B.L. | year=1993 | title=A Supplement to Distribution and Taxonomy of Birds of the World | place=New Haven, Connecticut | publisher=Yale University Press | isbn=978-0-300-05549-8 }}</ref><ref>{{ cite book | last1=Clement | first1=P. | last2=Hathway | first2=R. | year=2000 | title=Helm Identification Guides: Thrushes | place=London | publisher=Christopher Helm | isbn=978-07136-3940-7 | page=28 }}</ref> In 1998 the [[American Ornithologists' Union]] chose to treat the thrushes as a separate family in the seventh edition of their ''Check-list of North American birds'' and subsequently most authors have followed their example.<ref name=ioc>{{cite web| editor1-last=Gill | editor1-first=Frank | editor1-link=Frank Gill (ornithologist) | editor2-last=Donsker | editor2-first=David | editor3-last=Rasmussen | editor3-first=Pamela | editor3-link=Pamela C. Rasmussen | date=February 2025 | title=Chats, Old World flycatchers | work=IOC World Bird List Version 15.1 | url=https://www.worldbirdnames.org/bow/chats/ | publisher=International Ornithologists' Union | access-date=28 March 2025}}</ref><ref>{{cite book | author=Committee on Classification and Nomenclature | year=1998 | title=Check-list of North American birds | edition=7th | publisher=American Ornithologists' Union | place=Washington D.C. | isbn=1-891276-00-X | page=495 | url=http://americanornithology.org/sites/default/files/files/imce-img/AOUchecklistPass-Mimidae.pdf#80 | access-date=2016-07-08 | archive-url=https://web.archive.org/web/20151118161854/http://americanornithology.org/sites/default/files/files/imce-img/AOUchecklistPass-Mimidae.pdf#80 | archive-date=2015-11-18 | url-status=dead }}</ref> ===Genera=== The family formerly included fewer species. At the time of the publication of the third edition of [[Howard and Moore Complete Checklist of the Birds of the World]] in 2003, the genera ''[[Myophonus]]'', ''[[Alethe (genus)|Alethe]]'', ''[[Brachypteryx]]'' and ''[[Monticola]]'' were included in the thrush family [[Turdidae]].<ref name=h&m3>{{cite book | editor1-last=Dickinson | editor1-first=E.C. | year=2003 | title=The Howard and Moore Complete Checklist of the Birds of the World | edition=3rd | place=London | publisher=Christopher Helm | isbn=978-0-7136-6536-9 }}</ref> Subsequent [[molecular phylogenetic]] studies have shown that the species in these four genera are more closely related to species in Muscicapidae.<ref name=voelker2004>{{ cite journal | last1=Voelker | first1=G. | last2=Spellman | first2=G.M. | year=2004 | title=Nuclear and mitochondrial DNA evidence of polyphyly in the avian superfamily Muscicapoidea | journal=Molecular Phylogenetics and Evolution | volume=30 | issue= 2| pages=386–394 | doi=10.1016/S1055-7903(03)00191-X | pmid=14715230 | bibcode=2004MolPE..30..386V }}</ref><ref name=sangster2010>{{ cite journal | last1=Sangster | first1=G. | last2=Alström | first2=P. | last3=Forsmark | first3=E. | last4=Olsson | first4=U. | year=2010 | title=Multi-locus phylogenetic analysis of Old World chats and flycatchers reveals extensive paraphyly at family, subfamily and genus level (Aves: Muscicapidae) | journal=Molecular Phylogenetics and Evolution | volume=57 | issue=1 | pages=380–392 | doi=10.1016/j.ympev.2010.07.008 | pmid=20656044| bibcode=2010MolPE..57..380S }}</ref> As a consequence, these four genera are now placed here.<ref name=ioc/><ref name=h&m4>{{cite book | editor1-last=Dickinson | editor1-first=E.C. | editor2-last=Christidis | editor2-first=L. | year=2014 | title=The Howard & Moore Complete Checklist of the Birds of the World. | volume=2, Passerines | edition=4th | publisher=Aves Press | place=Eastbourne, U.K. | isbn=978-0-9568611-2-2 | pages=584, 598, 601, 607 }}</ref> In contrast, the genus ''[[Cochoa]]'' which was previously placed in Muscicapidae has been shown to belong in Turdidae.<ref name=voelker2004/><ref name=sangster2010/> Two large [[molecular phylogenetic]] studies of species within Muscicapidae published in 2010 showed that the genera ''Fraseria'', ''Melaenornis'' and ''Muscicapa'' were [[monophyletic|non-monophyletic]]. The authors were unable to propose revised genera as not all the species were sampled and not all the nodes in their phylogenies were strongly supported.<ref name=sangster2010/><ref>{{ cite journal | last1=Zuccon | first1=D. | last2=Ericson | first2=P.G.P. | year=2010 | title=A multi-gene phylogeny disentangles the chat-flycatcher complex (Aves: Muscicapidae) | journal=Zoologica Scripta | volume=39 | issue=3 | pages=213–224 | doi=10.1111/j.1463-6409.2010.00423.x | s2cid=85963319 }}</ref> A subsequent study published in 2016, that included 37 of the 42 Muscicapini species, confirmed that the genera were non-monophyletic and proposed a reorganised arrangement of the species with several new or resurrected genera.<ref>{{ cite journal | last1=Voelker | first1=G. | last2=Huntley | first2=J.W. | last3=Peñalba | first3=J.V. | last4=Bowie | first4=R.C.K. | year=2016 | title=Resolving taxonomic uncertainty and historical biogeographic patterns in ''Muscicapa'' flycatchers and their allies | journal=Molecular Phylogenetics and Evolution | volume=94 | issue= Pt B| pages=618–625 | doi=10.1016/j.ympev.2015.09.026 | pmid=26475615| doi-access=free | bibcode=2016MolPE..94..618V }}</ref> [[File:Madagascar magpie-robin (Copsychus albospecularis pica) female.jpg|thumb|Madagascar magpie-robin ''Copsychus albospecularis pica'']] [[File:Silverbird in Murchison Falls National Park, Uganda.JPG|thumb|right|[[Silverbird (bird)|Silverbird]], monotypic genus ''Empidornis'']] [[File:Tickell's Blue Flycatcher (Cyornis tickelliae) with feed W IMG 9340.jpg|thumb|[[Cyornis tickelliae|Tickell's blue flycatcher]], genus ''Cyornis'']] [[File:Amber mountain rock thrush (Monticola sharpei erythronotus) male 2.jpg|thumb|Amber mountain rock thrush ''Monticola sharpei erythronotus'']] [[File:Stonechat (Saxicola torquatus) male.jpg|thumb|European stonechat ''Saxicola torquatus'']] [[File:Cyprus wheatear (Oenanthe cypriaca).jpg|thumb|upright|Cyprus wheatear ''Oenanthe cypriaca'']] The [[International Ornithologists' Union]] recognises 357 species and divides the family into 57 genera. Subdivisions have been proposed by [[George Sangster|Sangster]] ''et al'' (2010).<ref name=ioc/><ref name=sangster2010/> For a complete list of species, see "[[List of Old World flycatcher species]]". '''Family Muscicapidae''' * '''Subfamily Muscicapinae''' ([[John Fleming (naturalist)|Fleming]], 1822) ** '''Tribe Copsychini''' (Sundevall, 1872) *** ''[[Alethe (genus)|Alethe]]'' – alethes *** ''[[Tychaedon]]'' – scrub robins *** ''[[Cercotrichas]]'' – scrub robins *** ''[[Copsychus]]'' – magpie-robins or shamas ** '''Tribe Muscicapini''' (Fleming, 1822) *** ''[[Agricola (bird)|Agricola]]'' *** ''[[Fraseria]]'' – forest flycatchers *** ''[[Melaenornis]]'' (7 species) *** ''[[Herero chat|Namibornis]]'' – single species: Herero chat *** ''[[Silverbird (bird)|Empidornis]]'' – single species: silverbird *** ''[[Sigelus]]'' – single species: fiscal flycatcher *** ''[[Bradornis]]'' (3 species) *** ''[[Myornis]]'' – single species: Böhm's flycatcher *** ''[[Artomyias]]'' (2 species) *** ''[[Humblotia (bird)|Humblotia]]'' – single species: Humblot's flycatcher *** ''[[Muscicapa]]'' (17 species) * '''Subfamily Niltavinae''' (Sangster, Alström, Forsmark and Olsson, 2010)<ref name=sangster2016>{{ cite journal | last1=Sangster | first1=G. | last2=Alström | first2=P. | last3=Forsmark | first3=Émile | last4=Olsson | first4=U. | year=2016 | title=Niltavinae, a new taxon of Old World flycatchers (Aves: Muscicapidae) | journal=Zootaxa | volume=4196 | issue=3 | pages=428–429 | doi=10.11646/zootaxa.4196.3.7 | pmid=27988667 | doi-access=free }}</ref>{{efn|The ornithologist Dario Zuccon pointed out that when George Sangster and colleagues erected the name "Niltavinae" for the subfamily, they did not provide a description as required by the [[International Code of Zoological Nomenclature]]. Sangster and colleagues subsequently published a description in 2016.<ref name=sangster2016/><ref name=zuccon2011>{{ cite journal | last=Zuccon | first=D. | year=2011 | title=Taxonomic notes on some Muscicapidae | journal=Bulletin of the British Ornithologists' Club | volume=131 | issue=3 | pages=196–199 | url=https://www.biodiversitylibrary.org/page/50781147 }}</ref>}} ** ''[[Leucoptilon]]'' – single species: white-tailed flycatcher ** ''[[Sholicola]]''<ref>{{cite journal |last1=Robin |first1=V.V. |last2=Vishnudas |first2=C. K. |last3=Gupta |first3=Pooja |last4=Rheindt |first4=Frank E. |last5=Hooper |first5=Daniel M. |last6=Ramakrishnan |first6=Uma |last7=Reddy |first7=Sushma |year=2017 |title=Two new genera of songbirds represent endemic radiations from the Shola Sky Islands of the Western Ghats, India |journal=BMC Evolutionary Biology |volume=17 |issue=1 |pages=31 |doi=10.1186/s12862-017-0882-6 |pmc=5259981 |pmid=28114902 |doi-access=free|bibcode=2017BMCEE..17...31R }}</ref> – sholakilis ** ''[[Niltava]]'' – niltavas ** ''[[Cyanoptila]]'' – flycatchers ** ''[[Eumyias]]'' – blue flycatchers ** ''[[Anthipes]]'' – flycatchers ** ''[[Cyornis]]'' – blue flycatchers * '''Subfamily Erithacinae''' ([[George Robert Gray|G.R. Gray]], 1846) – African forest robin assemblage{{efn|Dario Zuccon has argued that the correct name for the African forest robins assemblage is Cossyphinae (type genus ''Cossypha'' [[Nicholas Aylward Vigors|Vigors]], 1825) as the name predates Erithacinae (G.R. Gray, 1846).<ref name=zuccon2011/><ref>{{cite journal | last=Vigors | first=Nicholas Aylward | author-link=Nicholas Aylward Vigors | year=1825 | title=''Cossyphina'' | journal=Zoological Journal | volume=2 | page=395 | url=https://www.biodiversitylibrary.org/page/2255532}}</ref>}} ** ''[[Erithacus]]'' – single species: European robin ** ''[[Swynnertonia]]'' – single species: Swynnerton's robin ** ''[[Pogonocichla]]'' – single species: white-starred robin ** ''[[Stiphrornis]]'' – forest robins (3 species) ** ''[[Cossyphicula]]'' – robin-chats ** ''[[Chamaetylas]]'' – (4 species) ** ''[[Cossypha]]'' – robin-chats ** ''[[Cichladusa]]'' – palm thrushes ** ''[[Angola cave chat|Xenocopsychus]]'' – single species: Angola cave chat ** ''[[Dessonornis]]'' – robin-chats ** ''[[Sheppardia]]'' – akalats * '''Subfamily [[Saxicolinae]]''' ([[Nicholas Aylward Vigors|Vigors]], 1825) ** ''[[Irania (bird)|Irania]]'' – single species: white-throated robin ** ''[[Luscinia]]'' – nightingales and relatives ** ''[[Myiomela]]'' – robins ** ''[[Calliope (genus)|Calliope]]'' – rubythroats ** ''[[Enicurus]]'' – forktails ** ''[[Cinclidium (bird)|Cinclidium]]'' – single species: blue-fronted robin ** ''[[Myophonus]]'' – whistling thrushes ** ''[[Heinrichia]]'' – single species: great shortwing ** ''[[Vauriella]]'' ** ''[[Leonardina]]'' – single species: Bagobo babbler ** ''[[Brachypteryx]]'' – shortwings ** ''[[Larvivora]]'' – East and South-East Asian robins ** ''[[Ficedula]]'' – flycatchers ** ''[[Tarsiger]]'' – bush robins and bluetails ** ''[[Heteroxenicus]]'' – single species: Gould's shortwing ** ''[[Phoenicurus]]'' – redstarts ** ''[[Monticola]]'' – rock thrushes ** ''[[Saxicola]]'' – stonechats and chats ** ''[[Campicoloides]]'' – single species: buff-streaked chat ** ''[[Emarginata]]'' ** ''[[Pinarochroa]]'' – single species: moorland chat ** ''[[Thamnolaea]]'' – single species: mocking cliff chat ** ''[[Myrmecocichla]]'' – chats ** ''[[Oenanthe (bird)|Oenanthe]]'' – wheatears The [[cladogram]] below is based on a [[molecular phylogenetic]] study of the family by Min Zhao and collaborators that was published in 2023. Some regions of the phylogenetic tree were not strongly supported by the sequence data.<ref name=zhoa>{{Cite journal | last1=Zhao | first1=M. | last2=Gordon Burleigh | first2=J. | last3=Olsson | first3=U. | last4=Alström | first4=P. | last5=Kimball | first5=R.T. | date=2023 | title=A near-complete and time-calibrated phylogeny of the Old World flycatchers, robins and chats (Aves, Muscicapidae) | journal=Molecular Phylogenetics and Evolution | volume=178 | pages=107646 | doi=10.1016/j.ympev.2022.107646 | doi-access=free | pmid=36265831 | bibcode=2023MolPE.17807646Z }}</ref> Both the genera included and the number of species in each genera are taken from the list of birds maintained by [[Frank Gill (ornithologist)|Frank Gill]], [[Pamela C. Rasmussen]] and David Donsker on behalf of the [[International Ornithological Committee]] (IOC).<ref name=ioc/> {{Clade | style=font-size:90%;line-height:80% |label1=Muscicapidae |1={{Clade |label1=Muscicapinae |1={{clade |label1=Copsychini |1={{clade |1=''[[Alethe (bird)|Alethe]]'' – alethes (2 species) |2={{clade |1=''[[Tychaedon]]'' – scrub robins (5 species) |2={{clade |1=''[[Cercotrichas]]'' – scrub robins (5 species) |2=''[[Copsychus]]'' – magpie-robins, sharmas (17 species) }} }} }} |label2=Muscicapini |2={{clade |1=''[[Vauriella goodfellowi]]'' – slaty-backed jungle flycatcher (position uncertain) |2={{clade |1={{clade |1=''[[Agricola (bird)|Agricola]]'' – flycatchers (2 species) |2={{clade |1=''[[Fraseria]]'' – flycatchers (8 species) |2={{clade |1=''[[Melaenornis]]'' – flycatchers (7 species) |2={{clade |1=''[[Namibornis]]'' – herero chat |2={{clade |1=''[[Empidornis]]'' – silverbird |2=''[[Sigelus]]'' – fiscal flycatcher }} }} }} }} }} |2={{clade |1={{clade |1=''[[Bradornis]]'' – flycatchers (3 species) |2={{clade |1=''[[Myopornis]]'' – Böhm's flycatcher |2=''[[Artomyias]]'' – flycatchers (2 species) }} }} |2=''[[Muscicapa]]'' – flycatchers (17 species) & ''[[Humblotia (bird)|Humblotia]]'' – Humblot's flycatcher }} }} }} }} |2={{clade |label1=Niltavinae |1={{clade |1=''[[Leucoptilon]]'' – white-tailed flycatcher |2={{clade |1={{clade |1=''[[Sholicola]]'' – blue robins (2 species) |2={{clade |1=''[[Niltava]]'' – niltavas (7 species) |2={{clade |1=''[[Cyanoptila]]'' – flycatchers (2 species) |2=''[[Eumyias]]'' – flycatchers (6 species) }} }} }} |2={{clade |1=''[[Anthipes]]'' – flycatchers (2 species) |2=''[[Cyornis]]'' – blue flycatchers, jungle flycatchers (36 species) many taxa unsampled }} }} }} |2={{clade |label1=Cossyphinae |1={{clade |1=''[[Erithacus]]'' – European robin |2={{clade |1={{clade |1=''[[Swynnertonia]]'' – Swynnerton's robin |2=''[[Pogonocichla]]'' – white-starred robin }} |2={{clade |1={{clade |1=''[[Stiphrornis]]'' – forest robin (position uncertain) |2={{clade |1=''[[Cossyphicula]]'' – robin-chats (2 species) |2={{clade |1=''[[Chamaetylas]]'' – alethes (4 species) |2=''[[Cossypha]]'' - robin-chats (8 species) }} }} }} |2={{clade |1={{clade |1=''[[Cossypha cyanocampter]]'' – blue-shouldered robin-chat (position uncertain) |2={{clade |1=''[[Cichladusa]]'' – palm thrushes (3 species) |2={{clade |1=''[[Xenocopsychus]]'' – Angola cave chat |2=''[[Dessonornis]]'' – robin-chats, ground robins (4 species) }} }} }} |2=''[[Sheppardia]]'' – akalats (11 species) }} }} }} }} |label2=Saxicolinae |2={{clade |1={{clade |1={{clade |1={{clade |1=''[[Irania (bird)|Irania]]'' – white-throated robin |2=''[[Luscinia]]'' – nightingales, redstart, bluethroat (4 species) }} |2=''[[Calliope (genus)|Calliope]]'' – rubythroats (5 species) }} |2={{clade |1={{clade |1=''[[Enicurus]]'' – forktails (8 species) |2={{clade |1=''[[Cinclidium (bird)|Cinclidium]]'' – blue-fronted robin |2=''[[Myophonus]]'' – whistling thrushes (9 species) }} }} |2={{clade |1=''[[Myiomela]]'' – robins (3 species) |2={{clade |1={{clade |1={{clade |1=''[[Heinrichia]]'' – great shortwing |2=''[[Vauriella]]'' – jungle flycatchers (3 species) & ''[[Leonardina]] – ''Bagobo babbler }} |2={{clade |1=''[[Brachypteryx]]'' – shortwings (6 species) |2=''[[Larvivora]]'' – robins (6 species) }} }} |2=''[[Ficedula]]'' – flycatchers (34 species) }} }} }} }} |2={{clade |1=''[[Tarsiger]]'' – bush robins, bluetails (8 species) |2={{clade |1=''[[Heteroxenicus]]'' – Gould's shortwing |2={{clade |1=''[[Phoenicurus]]'' – redstarts (14 species) |2={{clade |1=''[[Monticola]]'' – rock thrushes (13 species) |2={{clade |1=''[[Saxicola]]'' – bush chats, stonechats (14 species) |2={{clade |1={{clade |1=''[[Campicoloides]]'' – buff-streaked chat |2=''[[Emarginata]]'' – chats (3 species) }} |2={{clade |1={{clade |1=''[[Pinarochroa]]'' – moorland chat |2=''[[Myrmecocichla]]'' – chats (8 species) & ''[[Thamnolaea]]'' – mocking cliff chat }} |2=''[[Oenanthe (bird)|Oenanthe]]'' – wheatears (32 species) }} }} }} }} }} }} }} }} }} }} }} }} ==Description== The appearance of these birds is very varied, but they mostly have weak songs and harsh calls. They are small to medium birds, ranging from 9 to 22 cm in length.<ref name="HBW11">del Hoyo, J.; Elliot, A. & Christie D. (editors). (2006). ''[[Handbook of the Birds of the World]]''. Volume 11: Old World Flycatchers to Old World Warblers. Lynx Edicions. {{ISBN|84-96553-06-X}}.</ref> Many species are dull brown in colour, but the [[plumage]] of some can be much brighter, especially in the males.<ref>{{Cite web |url=http://www.artfullbirds.com/Old%20World%20Flycatchers.html |title=Old World Flycatchers Muscicapidae |work=artfullbirds.com |access-date=June 3, 2010}}</ref> Most have broad, flattened bills suited to catching insects in flight, although the few ground-foraging species typically have finer bills.<ref name=EoB>{{cite book |last=Perrins |first=C. |editor-last=Forshaw |editor-first=Joseph |year=1991 |title=Encyclopaedia of Animals: Birds|publisher= Merehurst Press|location=London|pages= 194–195|isbn= 1-85391-186-0}}</ref> Old World flycatchers live in almost every environment with a suitable supply of trees, from dense forest to open scrub, and even the montane woodland of the [[Himalayas]]. The more northerly species migrate south in winter, ensuring a continuous diet of insects.<ref name=EoB/> Depending on the species, their [[bird nest|nest]]s are either well-constructed cups placed in a tree or cliff ledge, or simply lining in a pre-existing tree hole. The hole-nesting species tend to lay larger clutches, with an average of eight eggs, rather than just two to five.<ref name=EoB/> ==Notes== {{notelist}} ==References== {{Reflist}} ==External links== {{Commons|Muscicapidae}} * [http://ibc.lynxeds.com/family/tyrant-flycatchers-tyrannidae Old World flycatcher videos] on the Internet Bird Collection * {{Cite EB1911 |last=Newton |first=Alfred |authorlink=Alfred Newton |wstitle=Flycatcher |short=x}} * {{Cite Americana |last=Ingersoll |first=Ernest |authorlink=Ernest Ingersoll |wstitle=Flycatcher |short=x}} {{Taxonbar|from=Q200989}} {{Authority control}} [[Category:Muscicapidae|*]] [[Category:Taxa named by John Fleming (naturalist)]]
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