Editing Piculet

{{short description|Subfamily of birds}}
{{Automatic Taxobox
| name              = Piculets
| image             = Picumnus temminckii2.jpg
| image_caption     = [[Ochre-collared piculet]]
| taxon             = Picumninae
| subdivision_ranks = Genera
| subdivision       =
''[[Picumnus (bird)|Picumnus]]''<br>
''[[Verreauxia (bird)|Verreauxia]]''<br>
''[[Sasia]]''
}}

The '''piculets''' are a distinctive [[subfamily]], '''Picumninae''', of small [[woodpecker]]s which occur mainly in tropical [[South America]], with just three [[Asia]]n and one [[African piculet|African species]].

Like the true woodpeckers, piculets have large heads, long tongues which they use to extract their [[insect]] prey and [[zygodactyl]] feet, with two toes pointing forward, and two backwards. However, they lack the stiff tail feathers that the true woodpeckers use when climbing trees, so they are more likely than their relatives to perch on a branch rather than an upright trunk.

Their bills are shorter and less dagger-like than the true woodpeckers, so they look for insects and grubs mainly in decaying wood. Similarly, they re-use woodpecker holes for nesting, rather than making their own holes. The eggs are white, as with many hole nesters.

Typically these birds have grey or dull green upperparts and dark-streaked white underparts.

==Systematics and evolution==
Although not well known from [[fossils]], the evolution of piculets is now considered rather straightforward. The disjunct occurrence of the genera, with one African species of the Southeast Asian ''Sasia'' and one Southeast Asian species of the American ''Picumnus'' is of comparatively recent origin. Molecular dating, calibrated with geographic events in the absence of a good fossil record, points at the Late [[Miocene]], c. 8 [[mya (unit)|MYA]], as the point where the two genera divided into their two respective lineages. At that time, there was a notable [[global cooling]] period. The molecular distances between piculets and woodpeckers are comparatively small for subfamilies, agreeing with the hypothesis that the split between the three groups of woodpecker-like picids subfamilies occurred only during the [[Middle Miocene disruption|Miocene climatic optimum]], around 15 MYA. The later radiation of South American piculets is probably due to changes in [[topology]] and climate fluctuations during the [[Pliocene]] and [[Pleistocene]]. The genus ''Verreauxia'' may be accepted because of pronounced morphological similarities, but the two ''Picumnus'' lineages, despite having diverged long ago, are virtually alike except for head coloration.<ref>Fuchs ''et al.'', 2006.</ref>

The [[Antillean piculet]] (''Nesoctites micromegas'') has proven to be a very distinct species evolutionarily between piculets and woodpeckers<ref>Benz ''et al.'', 2006.</ref> and thus is nowadays placed in a subfamily of its own.<ref>{{cite journal | last1 = Benz | first1 = Brett W. | last2 = Robbins | first2 = Mark B. | last3 = Peterson | first3 = A. Townsend | year = 2006 | title = Evolutionary history of woodpeckers and allies (Aves: Picidae): Placing key taxa on the phylogenetic tree | journal = Molecular Phylogenetics and Evolution | pmid = 16635580 | volume = 40 | issue = 2| pages = 389–399 | doi = 10.1016/j.ympev.2006.02.021 }}</ref>

The arrangement of species in the genera is as follows<ref>del Hoyo ''et al.'' (2002)</ref>

Genus ''[[Picumnus (bird)|Picumnus]]''
* [[Speckled piculet]],  ''P. innominatus''
* [[Bar-breasted piculet]],  ''P. aurifrons''
* [[Orinoco piculet]],  ''P. pumilus''
* [[Lafresnaye's piculet]],  ''P. lafresnayi''
* [[Golden-spangled piculet]],  ''P. exilis''
* [[Black-dotted piculet]],  ''P. nigropunctatus''
* [[Ecuadorian piculet]],  ''P. sclateri''
* [[Scaled piculet]],  ''P. squamulatus''
* [[White-bellied piculet]],  ''P. spilogaster''
* [[Arrowhead piculet]],  ''P. minutissimus''
* [[Spotted piculet]],  ''P. pygmaeus''
* [[Speckle-chested piculet]],  ''P. steindachneri''
* [[Varzea piculet]],  ''P. varzeae''
* [[White-barred piculet]],  ''P. cirratus''
* [[Ocellated piculet]],  ''P. dorbignyanus''
* [[Ochre-collared piculet]],  ''P. temminckii''
* [[White-wedged piculet]],  ''P. albosquamatus''
* [[Rusty-necked piculet]],  ''P. fuscus''
* [[Rufous-breasted piculet]],  ''P. rufiventris''
* [[Ochraceous piculet]],  ''P. limae''
* [[Mottled piculet]],  ''P. nebulosus''
* [[Plain-breasted piculet]],  ''P. castelnau''
* [[Fine-barred piculet]],  ''P. subtilis''
* [[Olivaceous piculet]],  ''P. olivaceus''
* [[Greyish piculet]],  ''P. granadensis''<!-- BullBOC123:7 -->
* [[Chestnut piculet]],  ''P. cinnamomeus''

Genus ''[[Sasia]]''
* [[Rufous piculet]],  ''S. abnormis''
* [[White-browed piculet]],  ''S. ochracea''

Genus ''[[Verreauxia (bird)|Verreauxia]]''
* [[African piculet]], ''V. africana''

==References==
{{Reflist}}
* '''Benz''', Brett W.; Robbins, Mark B. & Peterson, A. Townsend (2006)  Evolutionary history of woodpeckers and allies (Aves: Picidae): Placing key taxa on the phylogenetic tree. ''Molecular Phylogenetics and Evolution'' '''40''': 389–399. {{doi|10.1016/j.ympev.2006.02.021}} {{PMID|16635580}} (HTML abstract)
* '''del Hoyo''', J.; Elliott, A. & Sargatal, J. (editors) (2002): ''[[Handbook of Birds of the World]], Volume 7: Jacamars to Woodpeckers''. Lynx Edicions, Barcelona. {{ISBN|84-87334-22-9}}
* '''Fuchs, J.'''; Ohlson, J. I.; Ericson, Per G. P. & Pasquet, E. (2006): Molecular phylogeny and biogeographic history of the piculets (Piciformes: Picumninae). ''Journal of Avian Biology'' '''37'''(5): 487–496. {{doi|10.1111/j.0908-8857.2006.03768.x}} (HTML abstract)

==External links==
*[http://ibc.lynxeds.com/family/woodpeckers-picidae Piculet videos, photos & sounds] on the Internet Bird Collection

{{Taxonbar|from=Q244364}}

[[Category:Picumninae| ]]